GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **376 - 400** of **1732** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▼	Monoisotopic Mass	Organisms
G03260OU	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	GM4 N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	3170.11	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ovis aries Ovis Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03277YI	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc(b1-	GM1b cisGM1	Linkage defined saccharide	998.34	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Pecora Phasianidae Phasianinae Pipidae Pipoidea Primates Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus cellular organisms
G03313QW	L-Gal(b1-6)GalNAc(?1-		Linkage defined saccharide	383.14	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03457IA	Neu5Ac(a2-8)Neu5Gc(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	Asialo-GM1 N-Glycolyl-GM2	Linkage defined saccharide	1305.43	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03542LT	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-4)]GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3170.11	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03021WA	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	1667.64	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03067VG	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	I' antigen Paragloboside Polysialic acid Type 2 LN2	Linkage defined saccharide	1363.48	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03361KC	Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Linkage defined saccharide	1014.34	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Pseudomonadota Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G02915XR	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Sialylated LDN	Linkage defined saccharide	2450.89	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03499BP	Gal(a1-3)Gal(b1-4)GlcNAc(b1-	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	545.2	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03681UF	GlcNAc(b1-2)[Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Type 1 LN	Linkage defined saccharide	2118.77	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04220HB	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1558.54	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Discoba Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Kinetoplastea Laurasiatheria Mammalia Metakinetoplastina Metazoa Opisthokonta Sarcopterygii Schizotrypanum Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G04109RU	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1583.57	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04445KN	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1275.46	Acer pseudoplatanus Acer Acereae Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicales Carica papaya Carica Caricaceae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Gunneridae Haplorrhini Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Magnoliopsida Mammalia Mesangiospermae Metazoa Neobatrachia Neognathae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Pecora Pentapetalae Phasianidae Phasianinae Primates Ranidae Ranoidea Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae asterids cellular organisms eudicotyledons lamiids malvids rosids
G04478DK	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-3)[GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	Blood group A (type 2) Blood group H (type 2) I' antigen	Linkage defined saccharide	2690.02	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05108UT	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	748.27	Aculeata Agelenidae Agelenopsis aperta Agelenopsis Alveolata Amniota Amphiesmenoptera Amphipyrinae Ananas comosus Ananas Anas platyrhynchos Anas sp. Anas Anatidae Anatinae Anseriformes Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arachnida Araneae Araneomorphae Archelosauria Archosauria Armoracia rusticana Armoracia Arthropoda Artiodactyla Ascomycota Aves Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicaceae Brassicales Bromeliaceae Bromelioideae Capra hircus Capra Caprinae Cardamineae Carica papaya Carica Caricaceae Carnivora Catarrhini Chelicerata Chordata Ciliophora Coelurosauria Coturnix coturnix Coturnix Craniata Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Ditrysia Ecdysozoa Embryophyta Endopterygota Entelegynae Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Hymenostomatida Insecta Intramacronucleata Komagataella pastoris Komagataella Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Oligohymenophorea Opisthokonta Ovis aries Ovis Panarthropoda Pancrustacea Pecora Pentapetalae Perdicinae Petrosaviidae Phaffomycetaceae Phasianidae Phasianinae Poales Primates Protostomia Pterygota RTA clade Rattus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sar Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrahymena pyriformis Tetrahymena Tetrahymenidae Tetrahymenina Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms commelinids eudicotyledons malvids rosids saccharomyceta unclassified Anas unclassified entries unclassified sequences unidentified
G04820ZT	Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1827.68	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04964UA	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	1972.71	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05187DJ	Fuc(a1-2)Gal(b1-3)GalNAc-ol		Linkage defined saccharide	531.22	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Opisthokonta Primates Rana dalmatina Rana Rana Ranidae Ranoidea Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05235KE	GalNAc(a1-3)[Neu5Ac(a2-6)]GalNAc-ol		Linkage defined saccharide	717.28	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05319BS	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic Polysialic acid	Linkage defined saccharide	1712.61	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05611SB	GalNAc4S(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	4O-sulfated LDN N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1907.63	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05724WC	GlcNAc(a1-4)Gal(b1-3)[GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-6)]GalNAc(?1-	N-acetyllactosamine O-Glycan core 2 fuzzy Type 2 LN	Linkage defined saccharide	1154.43	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05784QE	Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1115.42	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Digenea Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lophotrochozoa Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Platyhelminthes Primates Protostomia Ruminantia Sarcopterygii Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Simiiformes Spiralia Strigeidida Teleostomi Tetrapoda Theria Trematoda Vertebrata cellular organisms
G05855AL	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(?1-	Mannose N-acetyllactosamine O-Mannose Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms