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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **76 - 100** of **1500** in total

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GlyTouCan ID	IUPAC Condensed	Motifs ▼	Subsumption Level	Monoisotopic Mass	Organisms
G73051OD	Man(a1-3)[Man(a1-3)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1204.42	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Embryophyta Eukaryota Euphorbiaceae Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Malpighiales Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Ricinus communis Ricinus Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G80513FH	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1042.37	50 kb inversion clade Agaricaceae Agaricales Agaricineae Agaricomycetes Agaricomycetidae Agaricomycotina Agaricus bisporus Agaricus Amniota Archelosauria Archosauria Armoracia rusticana Armoracia Aves Basidiomycota Bilateria Brassicaceae Brassicales Cardamineae Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Embryophyta Eukaryota Eumetazoa Euphyllophyta Euteleostomi Fabaceae Fabales Fabeae Fungi Galliformes Galloanserae Gnathostomata Gunneridae Hologalegina IRL clade Lamiales Lophotrochozoa Magnoliopsida Mesangiospermae Metazoa Mollusca NPAAA clade Neognathae Nicotiana alata Nicotiana tabacum Nicotiana Nicotianeae Nicotianoideae Olea europaea Olea Oleaceae Oleeae Opisthokonta Papilionoideae Pentapetalae Perdicinae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pisum sativum Pisum Protostomia Sarcopterygii Sauria Saurischia Sauropsida Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum tuberosum Solanum Spermatophyta Spiralia Streptophyta Streptophytina Teleostomi Tetrapoda Theropoda Tracheophyta Vertebrata Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids malvids rosids
G83958MI	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1720.59	50 kb inversion clade Amniota Ascomycota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Komagataella pastoris Komagataella Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaffomycetaceae Phaseoleae Rodentia Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids saccharomyceta
G86191DF	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-6)]Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2044.7	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Glycine max Glycine subgen. Soja Glycine Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G90054TK	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1882.64	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Glycine max Glycine subgen. Soja Glycine Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G92612ZM	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1056.39	Acalypheae Acalyphoideae Aculeata Ambrosia artemisiifolia var. elatior Ambrosia artemisiifolia Ambrosia trifida Ambrosia Amphiesmenoptera Amphipyrinae Amygdaleae Amygdaloideae Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Asteraceae Asterales Asteroideae Bilateria Chloridoideae Cynodon dactylon Cynodon Cynodonteae Dicondylia Ditrysia Ecdysozoa Eleusininae Embryophyta Endopterygota Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Glossata Gunneridae Hadeninae Heliantheae alliance Heliantheae Heteroneura Hexapoda Hymenoptera Insecta Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mandibulata Mesangiospermae Metazoa Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta PACMAD clade Panarthropoda Pancrustacea Pentapetalae Petrosaviidae Poaceae Poales Protostomia Prunus dulcis Prunus Pterygota Ricinus communis Ricinus Rosaceae Rosales Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Tracheophyta Viridiplantae asterids campanulids cellular organisms commelinids eudicotyledons fabids lamiids rosids
G93516TF	Man(a1-2)Man(a1-3)[Man(a1-6)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1396.49	Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Pentapetalae Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G94692TX	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1391.51	50 kb inversion clade Acalypheae Acalyphoideae Acer pseudoplatanus Acer Acereae Chrysophylloideae Embryophyta Ericales Eukaryota Euphorbiaceae Euphyllophyta Fabaceae Fabales Gunneridae Hippocastanoideae Magnoliopsida Malpighiales Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Platanaceae Platanus Proteales Ricinus communis Ricinus Sapindaceae Sapindales Sapotaceae Spermatophyta Streptophyta Streptophytina Synsepalum dulcificum Synsepalum Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade malvids rosids
G95786NX	Man(a1-3)[Man(a1-6)]Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1396.49	Acalypheae Acalyphoideae Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Glires Gnathostomata Gunneridae Magnoliopsida Malpighiales Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pentapetalae Ricinus communis Ricinus Rodentia Sarcopterygii Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids rosids
G96906RY	Gal(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1072.38	Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Pentapetalae Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G28491QG	Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1245.45	BOP clade Embryophyta Eukaryota Euphyllophyta Liliopsida Magnoliopsida Mesangiospermae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Petrosaviidae Poaceae Poales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms commelinids
G45694ZE	Man(a1-3/6)[Man(a1-3/6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-?)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1188.43	Apiaceae Apiales Apiineae Apioideae Daucinae Daucus carota Daucus sect. Daucus Daucus Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Scandiceae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids campanulids cellular organisms eudicotyledons
G46273BF	Man(a1-3)[Man(a1-2)[Man(a1-3)]Man(a1-6)][Xyl(?1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1512.53	50 kb inversion clade Arachis hypogaea Arachis Dalbergieae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae Papilionoideae Pentapetalae Pterocarpus clade Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms dalbergioids sensu lato eudicotyledons fabids rosids
G49009JI	Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1391.51	50 kb inversion clade Brassica napus Brassica rapa subsp. chinensis Brassica rapa Brassica Brassicaceae Brassicales Brassiceae Caryophyllaceae Caryophyllales Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Silene latifolia subsp. alba Silene latifolia Silene sect. Melandrium Silene subgen. Behenantha Silene Sileneae Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids malvids rosids
G53401FX	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1234.43	50 kb inversion clade Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Gnathostomata Gunneridae Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muroidea Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Vigna radiata var. radiata Vigna radiata Vigna Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids saccharomyceta
G54833SQ	Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1245.45	Cucurbita pepo Cucurbita Cucurbitaceae Cucurbitales Cucurbiteae Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G60281CE	Gal(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1204.42	Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Pentapetalae Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G64979EW	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1042.37	50 kb inversion clade Acalypheae Acalyphoideae Amniota BOP clade Bilateria Boreoeutheria Chordata Craniata Cucurbitaceae Cucurbitales Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fabaceae Fabales Gastropoda Glires Gnathostomata Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Hygrophila Liliopsida Lophotrochozoa Lymnaea stagnalis Lymnaea Lymnaeidae Lymnaeoidea Magnoliopsida Malpighiales Mammalia Mesangiospermae Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panpulmonata Papilionoideae Pentapetalae Petrosaviidae Phaseoleae Phaseolus vulgaris Phaseolus Poaceae Poales Protostomia Ricinus communis Ricinus Rodentia Sarcopterygii Solanaceae Solanales Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae asterids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids rosids
G69195IF	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1245.45	Acer pseudoplatanus Acer Acereae BOP clade Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Lamiales Liliopsida Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Olea europaea Olea Oleaceae Oleeae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Pentapetalae Petrosaviidae Poaceae Poales Sapindaceae Sapindales Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms commelinids eudicotyledons lamiids malvids rosids
G70599KC	Gal(?1-4)Gal(?1-4)[Fuc(a1-3)]GlcNAc(?1-2)Man(a1-3)[Man(a1-2)Man(a1-6)][Xyl(?1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	2023.72	50 kb inversion clade Arachis hypogaea Arachis Dalbergieae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae Papilionoideae Pentapetalae Pterocarpus clade Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms dalbergioids sensu lato eudicotyledons fabids rosids
G75311GD	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1188.43	50 kb inversion clade Acalypheae Acalyphoideae Bilateria Caryophyllaceae Caryophyllales Embryophyta Erythrina corallodendron Erythrina Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euthyneura Fabaceae Fabales Gastropoda Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Magnoliopsida Malpighiales Mesangiospermae Metazoa Mollusca NPAAA clade Opisthokonta Panpulmonata Papilionoideae Pentapetalae Phaseoleae Protostomia Ricinus communis Ricinus Silene latifolia subsp. alba Silene latifolia Silene sect. Melandrium Silene subgen. Behenantha Silene Sileneae Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G79370VG	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1720.59	50 kb inversion clade Amniota Amylomyces rouxii Amylomyces Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi incertae sedis Fungi Glires Gnathostomata Gunneridae Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Mucoraceae Mucorales Mucorineae Mucoromycetes Mucoromycota Mucoromycotina Muridae Murinae Muroidea Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Rattus Rodentia Sarcopterygii Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G79754SY	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1720.59	50 kb inversion clade Amniota Ascomycota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Magnoliopsida Mammalia Mesangiospermae Metazoa Muroidea Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids saccharomyceta
G88330YM	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1245.45	BOP clade Embryophyta Eukaryota Euphyllophyta Gunneridae Liliopsida Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Pentapetalae Petrosaviidae Poaceae Poales Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms commelinids eudicotyledons lamiids
G35572VU	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic Type 1 LN	Linkage defined saccharide	1553.56	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids