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Repository for lectin-assisted multimodality data

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Tools

Glycan Converters

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Search Tools

Cross Search Glycan Search Publication Search Programmatic Access

Guidelines

MIRAGE

cytosolic thiouridylase subunit 2

Summary

Gene Symbol

CTU2

Organism

Homo sapiens (human)

External Links

NCBI Gene

348180

PubChem

348180

Alliance of Genome Resources

HGNC:28005

JoGo

CTU2

TogoVar

CTU2

Annotation

Keyword

Acetylation
Alternative splicing
Cytoplasm
Phosphoprotein
Primary microcephaly
Proteomics identification
Reference proteome
tRNA processing

Proteins

Displaying **all 2** entries
UniProt	Protein Name
H3BSW6
Q2VPK5	Cytosolic thiouridylase subunit 2

Annotation information from UniProt.

Gene Ontology (GO)

Displaying **all 4** entries
GO Term	Evidence Code	PMID
tRNA wobble uridine modification	NAS	19017811
tRNA wobble position uridine thiolation	IBA
protein urmylation	IEA
tRNA thio-modification	NAS	19017811

GO Hierarchy

biological process

metabolic process [inference]

nitrogen compound metabolic process [inference]

cellular nitrogen compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein modification process [inference]

post-translational protein modification [inference]

protein modification by small protein conjugation or removal [inference]

protein modification by small protein conjugation [inference]

protein urmylation

cellular metabolic process [inference]

cellular aromatic compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

cellular nitrogen compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

heterocycle metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

primary metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

protein metabolic process [inference]

protein modification process [inference]

post-translational protein modification [inference]

protein modification by small protein conjugation or removal [inference]

protein modification by small protein conjugation [inference]

protein urmylation

organic substance metabolic process [inference]

macromolecule metabolic process [inference]

protein metabolic process [inference]

protein modification process [inference]

post-translational protein modification [inference]

protein modification by small protein conjugation or removal [inference]

protein modification by small protein conjugation [inference]

protein urmylation

macromolecule modification [inference]

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

protein modification process [inference]

post-translational protein modification [inference]

protein modification by small protein conjugation or removal [inference]

protein modification by small protein conjugation [inference]

protein urmylation

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

organic cyclic compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein modification process [inference]

post-translational protein modification [inference]

protein modification by small protein conjugation or removal [inference]

protein modification by small protein conjugation [inference]

protein urmylation

cellular process [inference]

cellular metabolic process [inference]

cellular aromatic compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

cellular nitrogen compound metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

heterocycle metabolic process [inference]

nucleobase-containing compound metabolic process [inference]

nucleic acid metabolic process [inference]

RNA metabolic process [inference]

RNA processing [inference]

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

RNA modification [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA metabolic process [inference]

tRNA metabolic process [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

ncRNA processing [inference]

tRNA processing [inference]

tRNA modification [inference]

tRNA wobble base modification [inference]

tRNA wobble uridine modification

tRNA wobble position uridine thiolation

tRNA thio-modification

tRNA wobble position uridine thiolation

Displaying **all 3** entries
GO Term	Evidence Code	PMID
cytosol	TAS
cytosol	IBA
protein-containing complex	IDA	19017811

GO Hierarchy

cellular component

protein-containing complex

cellular anatomical entity [inference]

cytosol

Displaying **all 4** entries
GO Term	Evidence Code	PMID
tRNA binding	IEA
protein binding	IPI	19017811
nucleotidyltransferase activity	IEA
sulfurtransferase activity	IBA

GO Hierarchy

molecular function

catalytic activity [inference]

transferase activity [inference]

transferase activity, transferring phosphorus-containing groups [inference]

nucleotidyltransferase activity

transferase activity, transferring sulphur-containing groups [inference]

sulfurtransferase activity

binding [inference]

protein binding

organic cyclic compound binding [inference]

nucleic acid binding [inference]

RNA binding [inference]

tRNA binding

Gene Ontology information from NCBI Gene and UniProt.

OrthoDB (Group)

Group level

Eukaryota

Group Name

Cytoplasmic tRNA 2-thiolation protein 2 A

Functional Category

E: Amino acid transport and metabolism
G: Carbohydrate transport and metabolism
O: Posttranslational modification, protein turnover, chaperones

Displaying **all 7** entries
Gene Ontology
nucleotidyltransferase activity
protein urmylation
sulfurtransferase activity
tRNA processing
tRNA thio-modification
tRNA wobble position uridine thiolation
tRNA wobble uridine modification

Displaying **all 2** entries
InterPro
Cytoplasmic tRNA 2-thiolation protein 2
Rossmann-like alpha/beta/alpha sandwich fold

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GlyCosmos is a member of the GlySpace Alliance together with GlyGen and Glycomics@ExPASy.

Acknowledgements

Supported by JST NBDC Grant Number JPMJND2204

Partly supported by NIH Common Fund Grant #1U01GM125267-01

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GlyCosmos Portal v4.5.0

Last updated: April 6, 2026