RES
1r:r1
REP
REP1:4o(4+1)2d=-1--1
RES
2b:b-dgal-HEX-1:5
3s:n-acetyl
4b:b-dglc-HEX-1:5|6:a
5s:sulfate
LIN
1:2d(2+1)3n
2:2o(3+1)4d
3:2o(4+1)5n

• Grinnell F et al. Cellular adhesiveness and extracellular substrata. Int Rev Cytol. 1978;53:65-144. doi: 10.1016/s0074-7696(08)62241-x. PMID: 208994.

• Yada T et al. Occurrence of collagen and proteoglycan forms of type IX collagen in chick embryo cartilage. Production and characterization of a collagen form-specific antibody. J Biol Chem. 1992 May 5;267:(13):9391-7. PMID: 1577766.

• Aplin JD et al. Complex carbohydrates of the extracellular matrix structures, interactions and biological roles. Biochim Biophys Acta. 1982 Dec;694:(4):375-418. doi: 10.1016/0304-4157(82)90003-x. PMID: 6760897.

• Toyoda H et al. Structural analysis of glycosaminoglycans in Drosophila and Caenorhabditis elegans and demonstration that tout-velu, a Drosophila gene related to EXT tumor suppressors, affects heparan sulfate in vivo. J Biol Chem. 2000 Jan 28;275:(4):2269-75. doi: 10.1074/jbc.275.4.2269. PMID: 10644674.

• Christner JE et al. Immunological determinants of proteoglycans. Antibodies against the unsaturated oligosaccharide products of chondroitinase ABC-digested cartilage proteoglycans. J Biol Chem. 1980 Aug 10;255:(15):7102-5. PMID: 6156167.

• Yamamoto Y et al. Immunohistochemical localization of chondroitin sulfate in the forestomach of the sheep. Eur J Histochem. 1995;39:(4):265-72. PMID: 8835180.

• Nadanaka S et al. Chondroitin 4-O-sulfotransferase-1 modulates Wnt-3a signaling through control of E disaccharide expression of chondroitin sulfate. J Biol Chem. 2008 Oct 3;283:(40):27333-43. doi: 10.1074/jbc.M802997200. PMID: 18667431.

• Izumikawa T et al. Involvement of chondroitin sulfate synthase-3 (chondroitin synthase-2) in chondroitin polymerization through its interaction with chondroitin synthase-1 or chondroitin-polymerizing factor. Biochem J. 2007 May 1;403:(3):545-52. doi: 10.1042/BJ20061876. PMID: 17253960.

• Koike T et al. FAM20B is a kinase that phosphorylates xylose in the glycosaminoglycan-protein linkage region. Biochem J. 2009 Jun 26;421:(2):157-62. doi: 10.1042/BJ20090474. PMID: 19473117.

• Hay ED et al. Extracellular matrix. J Cell Biol. 1981 Dec;91:(3 Pt 2):205s-223s. doi: 10.1083/jcb.91.3.205s. PMID: 6172429.

• Kawashima H et al. Binding of a large chondroitin sulfate/dermatan sulfate proteoglycan, versican, to L-selectin, P-selectin, and CD44. J Biol Chem. 2000 Nov 10;275:(45):35448-56. doi: 10.1074/jbc.M003387200. PMID: 10950950.

• Oohira A et al. Developmentally regulated expression of a brain specific species of chondroitin sulfate proteoglycan, neurocan, identified with a monoclonal antibody IG2 in the rat cerebrum. Neuroscience. 1994 May;60:(1):145-57. doi: 10.1016/0306-4522(94)90210-0. PMID: 8052408.

• Izumikawa T et al. Identification of chondroitin sulfate glucuronyltransferase as chondroitin synthase-3 involved in chondroitin polymerization: chondroitin polymerization is achieved by multiple enzyme complexes consisting of chondroitin synthase family members. J Biol Chem. 2008 Apr 25;283:(17):11396-406. doi: 10.1074/jbc.M707549200. PMID: 18316376.

• Avnur Z et al. Spatial interrelationships between proteoglycans and extracellular matrix proteins in cell cultures. Exp Cell Res. 1985 Jun;158:(2):321-32. doi: 10.1016/0014-4827(85)90457-4. PMID: 3891385.