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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **4326 - 4350** of **9717** in total

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GlyTouCan ID	IUPAC Condensed ▲	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G13849YJ	GlcNAc(b1-2)Man(a1-3)[Man(a1-3/6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1421.52	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G41477DH	GlcNAc(b1-2)Man(a1-3)[Man(a1-3/6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	1423.53	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G17336FF	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1275.46	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G75896PD	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1275.46	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G42680EX	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1113.41	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Glires Gnathostomata Gunneridae Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Pecora Pentapetalae Rodentia Ruminantia Sarcopterygii Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae asterids cellular organisms eudicotyledons lamiids
G70073SG	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1113.41	Acer pseudoplatanus Acer Acereae Achelata Alpharhabdovirinae Amniota Amphibia Amphiesmenoptera Amphipyrinae Anas sp. Anas Anatidae Anatinae Anseriformes Anura Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crustacea Decapoda Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Euarchontoglires Eucarida Eukaryota Eumalacostraca Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Magnoliopsida Malacostraca Mammalia Mandibulata Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Orthornavirae Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pecora Pentapetalae Perdicinae Phasianidae Phasianinae Pleocyemata Plusiinae Primates Protostomia Pterygota Ranidae Ranoidea Rhabdoviridae Riboviria Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids cellular organisms eudicotyledons lamiids malvids rosids unclassified Anas unclassified Vesiculovirus
G61751GZ	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1113.41	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G09362PR	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1115.42	Acer pseudoplatanus Acer Acereae Achelata Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Arthropoda Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Crustacea Decapoda Deuterostomia Dinosauria Dipnotetrapodomorpha Ecdysozoa Embryophyta Euarchontoglires Eucarida Eukaryota Eumalacostraca Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gnathostomata Gunneridae Haplorrhini Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Magnoliopsida Malacostraca Mammalia Mandibulata Mesangiospermae Metazoa Multicrustacea Neognathae Opisthokonta Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pentapetalae Platanaceae Platanus Pleocyemata Primates Proteales Protostomia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids
G64518MK	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)][Fuc(a1-6)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1405.52	Amphiesmenoptera Amphipyrinae Arthropoda Bilateria Dicondylia Ditrysia Ecdysozoa Endopterygota Eukaryota Eumetazoa Glossata Hadeninae Heteroneura Hexapoda Insecta Lepidoptera Mamestra brassicae Mamestra Mandibulata Metazoa Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Spodoptera frugiperda Spodoptera cellular organisms
G31416KB	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)][Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1405.52	Amphiesmenoptera Amphipyrinae Arthropoda Bilateria Dicondylia Ditrysia Ecdysozoa Endopterygota Eukaryota Eumetazoa Glossata Hadeninae Heteroneura Hexapoda Insecta Lepidoptera Mamestra brassicae Mamestra Mandibulata Metazoa Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Spodoptera frugiperda Spodoptera cellular organisms
G12593TX	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1259.47	Acrididae Acrididea Acridoidea Acridomorpha Amniota Amphiesmenoptera Amphipyrinae Arthropoda Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Caelifera Catarrhini Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lepidoptera Locusta migratoria Locusta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Oedipodinae Opisthokonta Orthoptera Panarthropoda Pancrustacea Polyneoptera Primates Protostomia Pterygota Rodentia Sarcopterygii Simiiformes Spodoptera frugiperda Spodoptera Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G14576KZ	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1259.47	Acrididae Acrididea Acridoidea Acridomorpha Amniota Amphiesmenoptera Arthropoda Artiodactyla Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Caelifera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Endopterygota Enoplea Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Laurasiatheria Lepidoptera Locusta migratoria Locusta Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nematoda Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Oedipodinae Opisthokonta Orthoptera Ovis aries Ovis Panarthropoda Pancrustacea Pecora Perissodactyla Plusiinae Polyneoptera Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata cellular organisms
G07483YN	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1259.47	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G27314YB	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	1261.48	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G49889OJ	GlcNAc(b1-2)Man(a1-3)[Man(a1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1275.46	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03597FX	GlcNAc(b1-2)Man(a1-3)[Man(a1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1421.52	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86795EH	GlcNAc(b1-2)Man(a1-3)[Man6S(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	1193.36	Achelata Arthropoda Bilateria Crustacea Decapoda Ecdysozoa Eucarida Eukaryota Eumalacostraca Eumetazoa Malacostraca Mandibulata Metazoa Multicrustacea Opisthokonta Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pleocyemata Protostomia cellular organisms
G97217ZC	GlcNAc(b1-2)Man(a1-3)[Man6S(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1195.38	Achelata Arthropoda Bilateria Crustacea Decapoda Ecdysozoa Eucarida Eukaryota Eumalacostraca Eumetazoa Malacostraca Mandibulata Metazoa Multicrustacea Opisthokonta Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pleocyemata Protostomia cellular organisms
G48489PM	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(?2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1915.69	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63787RZ	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	1915.69	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89643QK	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1566.56	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G82695BC	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(?1-	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1566.56	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70851FQ	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1769.63	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G90725ZC	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1769.63	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19447ZX	GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1915.69	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms