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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **26 - 50** of **666** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G30888FF	Glc(?1-2)Glc(?1-3)Glc(?1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1720.59	Alpharhabdovirinae Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Rhabdoviridae Riboviria Vesicular stomatitis virus Vesiculovirus Viruses unclassified Vesiculovirus
G31429YW	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan high mannose	Linkage defined saccharide	1720.59	Acalypheae Acalyphoideae Amniota Amphiesmenoptera Arthropoda Artiodactyla Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Embryophyta Endopterygota Euarchontoglires Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Orthoparamyxovirinae Orthornavirae Panarthropoda Pancrustacea Paramyxoviridae Pecora Pentapetalae Primates Protostomia Pterygota Respirovirus muris Respirovirus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids rosids
G35748HH	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3681.3	Amniota Artverviricota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend spleen focus-forming virus Gammaretrovirus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Gammaretrovirus
G36280LM	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	3074.07	Artverviricota Human immunodeficiency virus Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Retroviridae Revtraviricetes Riboviria Viruses unclassified Primate lentivirus group
G38271CW	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Lewis x Lex N-Glycan core basic N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1567.58	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G40452ID		N-Glycan core basic	Linkage defined saccharide	1113.41	Amniota Artverviricota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Human immunodeficiency virus Lentivirus Mammalia Metazoa Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G43710ZF	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[D-Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2315.85	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G44210YC	Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[D-Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2315.85	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G00031MO	Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	383.14	Acalyptratae Acanthomorphata Actinopteri Actinopterygii Amniota Arthropoda Artiodactyla Bacteria Bacteroidaceae Bacteroidales Bacteroides ovatus Bacteroides Bacteroidia Bacteroidota/Chlorobiota group Bacteroidota Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria FCB group Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Neopterygii Neoteleostei Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Panarthropoda Pancrustacea Paracanthopterygii Pecora Pisoniviricetes Pisuviricota Pollachius pollachius Pollachius Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Shigella dysenteriae Shigella Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses Zeiogadaria cellular organisms melanogaster group melanogaster subgroup
G00047MO	Gal(b1-3)[Fuc(a1-4)]GlcNAc(?1-	Lea Lewis a	Linkage defined saccharide	529.2	Amniota Artverviricota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Lentivirus Mammalia Metazoa Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Primates Retroviridae Revtraviricetes Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G00052MO	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-	Blood group H (type 2) Leb Lewis b Ley	Linkage defined saccharide	675.26	Amniota Artiodactyla Artverviricota Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Helicobacter pylori 26695 Helicobacter pylori SS1 Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Lagomorpha Laurasiatheria Lentivirus Leporidae Mammalia Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Oryctolagus cuniculus Oryctolagus Pararnavirae Pecora Primate lentivirus group Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G00033MO	Gal(b1-3)[GlcNAc(b1-6)]GalNAc(a1-	O-Glycan core 2	Linkage defined saccharide	586.22	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G00194GV			Base composition with linkage	367.15	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G00482HJ	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage defined saccharide	709.26	Alphaherpesvirinae Amniota Archelosauria Archosauria Articulavirales Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Duplodnaviria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fowl plague virus Galliformes Galloanserae Gnathostomata Haplorrhini Herpes simplex virus unknown type Herpesvirales Herviviricetes Heunggongvirae Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Negarnaviricota Neognathae Opisthokonta Orthoherpesviridae Orthomyxoviridae Orthornavirae Peploviricota Polyploviricotina Primates Riboviria Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Simplexvirus Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unclassified Simplexvirus unidentified influenza virus
G00406II			Glycosidic topology	1218.44	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G01169EW	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1786.65	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G01999JH	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(a1-	N-Glycan hybrid	Linkage defined saccharide	1437.51	Alpharhabdovirinae Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Rhabdoviridae Riboviria Vesicular stomatitis virus Vesiculovirus Viruses unclassified Vesiculovirus
G02633ON	GlcNAc(b1-2)Man(a1-6)Man(a1-3)Man(a1-6)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1439.53	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G02315DX			Glycosidic topology	2499.9	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G02628JF			Glycosidic topology	2305.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G02030ZB			Glycosidic topology	2280.82	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G02402FF		N-Glycan core basic	Linkage defined saccharide	1072.38	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G02815KT			Base composition with linkage	1234.43	Amarillovirales Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Flasuviricetes Flaviviridae Glires Gnathostomata Haplorrhini Hepacivirus hominis Hepacivirus Hepatitis C virus (isolate H77) Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms hepatitis C virus genotype 1a
G03382KH			Glycosidic topology	1827.68	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G03127AL		N-Glycan core basic	Linkage defined saccharide	1259.47	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms