GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **26 - 50** of 92 in total

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GlyTouCan ID	IUPAC Condensed ▼	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G86053IC	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1558.54	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Amniota Amphiesmenoptera Amphipyrinae Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Aves BOP clade Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Brassicales Carica papaya Carica Caricaceae Carnivora Carpinus betulus Carpinus Catarrhini Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Crithidia fasciculata Crithidia Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo IRL clade Insecta Insthoviricetes Kinetoplastea Lamiales Laurasiatheria Leishmaniinae Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Phasianinae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses asterids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G70358QM	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1722.61	50 kb inversion clade Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Kinetoplastea Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Neoptera Opisthokonta Panarthropoda Pancrustacea Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Primates Protostomia Pterygota Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids
G59513YA	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Acalypheae Acalyphoideae Acalyptratae Amniota Amphiesmenoptera Arthropoda Articulavirales Artiodactyla Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Insthoviricetes Kinetoplastea Laurasiatheria Lepidoptera Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unidentified influenza virus
G84170WO	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1722.61	50 kb inversion clade Acalyptratae Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Ascomycota Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Coelurosauria Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Galliformes Galloanserae Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Influenza A virus Insecta Insthoviricetes Kinetoplastea Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neoptera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unidentified influenza virus
G86385RC	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Alpharhabdovirinae Amniota Amphiesmenoptera Amphipyrinae Anserineae Anthemideae Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae Aves BOP clade Batoidea Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Carpinus betulus Carpinus Caryophyllales Catarrhini Cavia porcellus Cavia Caviidae Chenopodiaceae Chenopodioideae Chondrichthyes Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Fungi Galliformes Galloanserae Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha IRL clade Insecta Insthoviricetes Kinetoplastea Komagataella pastoris Komagataella Lamiales Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaffomycetaceae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Rhabdoviridae Riboviria Ricinus communis Ricinus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Schizotrypanum Simiiformes Sophophora Spermatophyta Spinacia oleracea Spinacia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified Vesiculovirus unidentified influenza virus
G66257KQ	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1884.66	50 kb inversion clade Acalyptratae Aculeata Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Articulavirales Artiodactyla Artverviricota Bilateria Boreoeutheria Bovidae Brachycera Caprinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fabeae Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Haplorrhini Hexapoda Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Hymenoptera IRL clade Influenza A virus Insecta Insthoviricetes Kinetoplastea Laurasiatheria Lens culinaris Lens Lentivirus Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neoptera Opisthokonta Ortervirales Orthomyxoviridae Orthoretrovirinae Orthornavirae Ovis aries Ovis Panarthropoda Pancrustacea Papilionoideae Pararnavirae Pecora Pentapetalae Phaseoleae Phaseolus coccineus Phaseolus lunatus Phaseolus vulgaris Phaseolus Polyploviricotina Primate lentivirus group Primates Protostomia Pterygota Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unclassified Primate lentivirus group unidentified influenza virus
G60230HH	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1882.64	50 kb inversion clade Acalyptratae Amniota Arachnida Arthropoda Ascomycota Bilateria Boreoeutheria Brachycera Catarrhini Chelicerata Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Neoptera Opisthokonta Panarthropoda Pancrustacea Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Schizophora Scorpiones Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids saccharomyceta
G10471FG	Man(?1-?)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1396.49	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G42666HT	GlcNAc(b1-4)GlcNAc(b1-		Linkage defined saccharide	424.17	Acalyptratae Amniota Arthropoda Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muscomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Primates Protostomia Pseudomonadota Pterygota Ruminantia Sarcopterygii Schizophora Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G07483YN	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1259.47	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G80393PG	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1665.62	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G80858MF	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1462.54	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Craniata Cricetidae Cricetinae Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G39213VZ	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1316.49	Acalyptratae Amniota Archelosauria Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Coelurosauria Craniata Cyclorrhapha Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G49108TO	GlcNAc(b1-				Acalyptratae Amniota Arthropoda Articulavirales Artiodactyla Ascomycota Bacilli Bacillota Bacteria Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Insthoviricetes Lactobacillales Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Negarnaviricota Neoptera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Polyploviricotina Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Schizophora Simiiformes Sophophora Streptococcaceae Streptococcus pneumoniae Streptococcus Suidae Suina Sus scrofa Sus Teleostomi Terrabacteria group Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup saccharomyceta unclassified Orthomyxoviridae unidentified influenza virus
G89319AW	GlcNAc(?1-?)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1437.51	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G39943KJ	GlcNAc(?1-?)Man(a1-3)[GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1316.49	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G54406UD	GlcA3S(b1-3)Gal(b1-4)GlcNAc(?1-		Linkage defined saccharide	639.13	Acalyptratae Amniota Archelosauria Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Coelurosauria Craniata Cyclorrhapha Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Phasianidae Phasianinae Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms melanogaster group melanogaster subgroup
G57321FI	GalNAc(a1-	Tn antigen			Acalyptratae Amniota Armadillidae Armadillo Arthropoda Artiodactyla Artverviricota Ascaridoidea Ascaridomorpha Ascomycota Bacteria Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenorhabditis elegans Caenorhabditis Calliphoridae Calyptratae Caprinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cestoda Chlorocebus aethiops Chlorocebus Chordata Chromadorea Chrysomyinae Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crinocheta Crustacea Cyclophyllidea Cyclorrhapha Deuterostomia Dicondylia Digenea Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Echinococcus granulosus group Echinococcus granulosus Echinococcus Echinostomata Echinostomatoidea Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eucestoda Eukaryota Eumalacostraca Eumetazoa Euteleostomi Eutheria Fasciola hepatica Fasciola Fasciolidae Fungi Gammaproteobacteria Glires Gnathostomata Haplorrhini Heligmosomidae Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Insecta Isopoda Lagomorpha Laurasiatheria Lentivirus Leporidae Lophotrochozoa Macaca mulatta Macaca Malacostraca Mammalia Mandibulata Mesocestoides corti Mesocestoides vogae Mesocestoides Mesocestoididae Metazoa Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Nematoda Neoptera Nidovirales Nippostrongylus brasiliensis Nippostrongylus Oestroidea Oniscidea Opisthokonta Ortervirales Orthocoronavirinae Orthoretrovirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Panarthropoda Pancrustacea Pararnavirae Pecora Peloderinae Peracarida Phormini Pisoniviricetes Pisuviricota Plagiorchiida Platyhelminthes Primate lentivirus group Primates Protophormia terraenovae Protophormia Protostomia Pseudomonadaceae Pseudomonadales Pseudomonadota Pseudomonas aeruginosa group Pseudomonas aeruginosa Pseudomonas Pterygota Rattus norvegicus Rattus Retroviridae Revtraviricetes Rhabditida Rhabditidae Rhabditina Rhabditoidea Rhabditomorpha Riboviria Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Spiralia Spirurina Strongyloidea Taenia hydatigena Taenia Taeniidae Teleostomi Tetrapoda Theria Toxocara canis Toxocara Toxocaridae Trematoda Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup saccharomyceta unclassified Primate lentivirus group
G00033MO	Gal(b1-3)[GlcNAc(b1-6)]GalNAc(a1-	O-Glycan core 2	Linkage defined saccharide	586.22	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G00031MO	Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	383.14	Acalyptratae Acanthomorphata Actinopteri Actinopterygii Amniota Arthropoda Artiodactyla Bacteria Bacteroidaceae Bacteroidales Bacteroides ovatus Bacteroides Bacteroidia Bacteroidota/Chlorobiota group Bacteroidota Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria FCB group Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Neopterygii Neoteleostei Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Panarthropoda Pancrustacea Paracanthopterygii Pecora Pisoniviricetes Pisuviricota Pollachius pollachius Pollachius Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Shigella dysenteriae Shigella Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses Zeiogadaria cellular organisms melanogaster group melanogaster subgroup
G19059PI					Acalyptratae Actinopteri Actinopterygii Amniota Ampullariidae Ampullarioidea Archelosauria Architaenioglossa Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenogastropoda Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Gerbillinae Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Lagomorpha Laurasiatheria Leporidae Lophotrochozoa Macaca mulatta Macaca Mammalia Mandibulata Meriones unguiculatus Meriones Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Pomacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spiralia Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Whippomorpha cellular organisms melanogaster group melanogaster subgroup
G43702JT					Acalyptratae Amniota Arthropoda Bacteria Bilateria Boreoeutheria Brachycera Caenorhabditis elegans Caenorhabditis Catarrhini Chordata Chromadorea Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Leporidae Mammalia Mandibulata Metazoa Moraxella bovis Moraxella Moraxellaceae Moraxellales Muridae Murinae Muroidea Muscomorpha Myomorpha Nematoda Neoptera Opisthokonta Oryctolagus cuniculus Oryctolagus Panarthropoda Pancrustacea Pasteurella multocida Pasteurella Pasteurellaceae Pasteurellales Peloderinae Primates Protostomia Pseudomonadota Pterygota Rattus Rhabditida Rhabditidae Rhabditina Rhabditoidea Rhabditomorpha Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G00406II			Glycosidic topology	1218.44	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G02402FF		N-Glycan core basic	Linkage defined saccharide	1072.38	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G03842KQ			Glycosidic topology	1907.68	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup