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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **501 - 525** of **583** in total

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GlyTouCan ID ▼	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G13974DV	Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1236.45	Acalypheae Acalyphoideae Acalyptratae Achelata Aculeata Amniota Anthophila Apidae Apocrita Apoidea Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Coelurosauria Craniata Crustacea Cyclorrhapha Decapoda Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Eucarida Eukaryota Eumalacostraca Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Insthoviricetes Lagomorpha Laurasiatheria Leporidae Magnoliopsida Malacostraca Malpighiales Mammalia Mandibulata Mesangiospermae Metazoa Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Negarnaviricota Neognathae Neoptera Opisthokonta Orthomyxoviridae Orthornavirae Oryctolagus cuniculus Oryctolagus Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pecora Pentapetalae Phasianidae Phasianinae Pleocyemata Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unidentified influenza virus
G13869TL	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1560.55	50 kb inversion clade Acalyptratae Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Coelurosauria Craniata Cyclorrhapha Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Influenza A virus Insecta Insthoviricetes Kinetoplastea Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neoptera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Phasianinae Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unidentified influenza virus
G13751GY	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-3/4)GlcNAc-ol		Linkage defined saccharide	1681.58	Alpharhabdovirinae Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Rhabdoviridae Riboviria Vesicular stomatitis virus Vesiculovirus Viruses unclassified Vesiculovirus
G13746SQ	Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	2206.75	Alpharhabdovirinae Amniota Archelosauria Archosauria Artiodactyla Asterias rubens Asterias Asteriidae Asteroidea Asterozoa Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Coelurosauria Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Echinodermata Eleutherozoa Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Forcipulatacea Forcipulatida Galliformes Galloanserae Glires Gnathostomata Haploviricotina Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Negarnaviricota Neognathae Opisthokonta Orthornavirae Pecora Perdicinae Phasianidae Rhabdoviridae Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata Vesicular stomatitis virus Vesiculovirus Viruses cellular organisms unclassified Vesiculovirus unclassified entries unclassified sequences unidentified
G13728QT			Glycosidic topology	2604.93	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G13716SG			Base composition with linkage	3612.31	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G13583BZ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Archelosauria Archosauria Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Laurasiatheria Mammalia Metazoa Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Phasianidae Phasianinae Polyploviricotina Primates Riboviria Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G13193DT			Base composition with linkage	2777.02	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G12872WY			Base composition with linkage	3815.39	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G12849CJ			Glycosidic topology	1761.62	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G12793SR			Glycosidic topology	3244.14	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G12580WI			Monosaccharide composition with linkage	2280.82	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G12341GU			Monosaccharide composition with linkage	3027.08	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G12313PD			Glycosidic topology	1729.63	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G11911BT			Glycosidic topology	2176.81	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G11870QZ			Glycosidic topology	1712.61	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G11700UP	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2516.91	Alphavirus Alsuviricetes Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Martellivirales Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Orthornavirae Primates Rattus Riboviria Rodentia Sarcopterygii Semliki Forest virus Simiiformes Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms
G11629QQ			Glycosidic topology	2733.97	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G11460AB	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)][Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1202.44	Acalyptratae Aculeata Amniota Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G11115RO			Monosaccharide composition with linkage	3535.24	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G10927TS	Man(a1-2)Man(a1-2)Man(a1-3)[Man(b1-3)[Man(a1-6)]Glc(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	1558.54	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G10773YW			Glycosidic topology	2036.72	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G10529ZR	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 1 LN Type 2 LN	Linkage defined saccharide	1932.71	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G10486CT			Base composition with linkage	1640.59	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G10374FO	GlcNAc(?1-?)Man(a1-3)[GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-?)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1462.54	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms