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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **5576 - 5600** of **8356** in total

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GlyTouCan ID	IUPAC Condensed ▲	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G79389NT	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM2	Linkage defined saccharide	836.29	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Carassius auratus Carassius Catarrhini Chordata Clupeocephala Craniata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Protacanthopterygii Salmonidae Salmoniformes Salmoninae Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G37481QP	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(?1-	3-Sialyl-LN (type 2)	Linkage defined saccharide	877.32	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G41052GC	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-	Sda/CT antigen	Linkage defined saccharide	877.32	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G50135PA	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Sda/CT antigen	Linkage defined saccharide	2775.0	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G50508SG	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex Sda/CT antigen	Linkage defined saccharide	4493.61	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G40343KH	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Sda/CT antigen	Linkage defined saccharide	4493.61	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G57835AK	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Sda/CT antigen	Linkage defined saccharide	4493.61	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G91846CD	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-3)Gal	Sda/CT antigen	Linkage defined saccharide	1039.37	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G83633HB	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-3)Gal(?1-	Sda/CT antigen	Linkage defined saccharide	1039.37	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G85563LA	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-3)Gal-ol	Sda/CT antigen	Linkage defined saccharide	1041.39	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89393WA	Neu5Ac(a2-3)[GalNAc(b1-?)[Neu5Ac(a2-3)]Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1a	Linkage defined saccharide	1492.52	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G21518JT	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc	GM4	Linkage defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Zeiogadaria cellular organisms
G46677TE	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1a	Linkage defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carangaria Carassius auratus Carassius Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chinchillidae Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Delphinidae Deuterostomia Dinosauria Dipnotetrapodomorpha Discoba Euacanthomorphacea Euarchontoglires Euglenozoa Eukaryota Eulipotyphla Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Kinetoplastea Lagomorpha Laurasiatheria Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Leporidae Mammalia Meleagridinae Meleagris gallopavo Meleagris Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Odontoceti Oncorhynchus mykiss Oncorhynchus Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Papio hamadryas Papio Parophrys vetulus Parophrys Pecora Percomorphaceae Phasianidae Phasianinae Pleuronectidae Pleuronectiformes Pleuronectoidei Primates Protacanthopterygii Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Soricidae Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trypanosomatida Trypanosomatidae Tursiops truncatus Tursiops Vertebrata Whippomorpha cellular organisms
G02149VR	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1aa	Linkage defined saccharide	1580.53	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pecora Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Selachii Squalidae Squaliformes Squalomorphii Squalus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G94278FQ	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-4)]GalNAc(b1-3)Gal(b1-3)Glc(b1-	GM4 Glucosylceramide	Linkage defined saccharide	1289.44	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G56577TY	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-4)]GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2659.94	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03542LT	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-4)]GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3170.11	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G92175CL	Neu5Ac(a2-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1a	Linkage defined saccharide	1492.52	Amniota Bilateria Boreoeutheria Caniformia Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Otariidae Pinnipedia Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G72769FF	Neu5Ac(a2-3)[Neu5Ac(a2-6)Gal(b1-4)]GlcNAc(b1-2)Man(a1-?)[Neu5Ac(a2-6)Gal(b1-?)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	2513.88	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03433NI	Neu5Ac(a2-3)[Neu5Ac(a2-6)]Gal(b1-3)GalNAc	GM4 O-Glycan core 1 fuzzy	Linkage defined saccharide	965.33	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70265CT	Neu5Ac(a2-3)[Neu5Ac(a2-6)]Gal(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc(b1-	GL-7 globoseries ganglioside SSEA-4	Linkage defined saccharide	1451.49	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G47867DI	Neu5Ac(a2-3)[Neu5Ac(a2-6)]Gal(b1-3)GalNAc-ol	GM4	Linkage defined saccharide	967.35	Amniota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muroidea Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84908OR	Neu5Ac(a2-3)[Neu5Ac(a2-6)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid a-Gal antigen	Linkage defined saccharide	2530.89	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G94019IV	Neu5Ac(a2-3)[Neu5Ac(a2-6)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2659.94	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G68110IF	Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1a	Linkage defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Bilateria Boreoeutheria Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Catarrhini Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Epsilonproteobacteria Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Primates Pseudocrenilabrinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms