GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **51 - 75** of **583** in total

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GlyTouCan ID	IUPAC Condensed ▲	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G13583BZ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Archelosauria Archosauria Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Laurasiatheria Mammalia Metazoa Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Phasianidae Phasianinae Polyploviricotina Primates Riboviria Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G32371XG	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1991.75	Amniota Articulavirales Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Polyploviricotina Primates Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G56791SN	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2354.86	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthomarburgvirus marburgense Orthomarburgvirus Orthoretrovirinae Orthornavirae Pararnavirae Pecora Perissodactyla Primate lentivirus group Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G83433HW	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2356.88	Amniota Artverviricota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Marburgvirus Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthoretrovirinae Orthornavirae Pararnavirae Perissodactyla Primate lentivirus group Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G35679TE	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2719.99	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haploviricotina Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthomarburgvirus marburgense Orthomarburgvirus Orthoretrovirinae Orthornavirae Pararnavirae Pecora Perissodactyla Primate lentivirus group Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G44325XR	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2722.01	Amniota Artverviricota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haploviricotina Human immunodeficiency virus 1 Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Marburgvirus Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthoretrovirinae Orthornavirae Pararnavirae Perissodactyla Primate lentivirus group Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G03740OH	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1786.65	Amniota Articulavirales Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Muroidea Myomorpha Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Polyploviricotina Primates Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G06301TU	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1788.67	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G62973JK	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2297.84	Amniota Articulavirales Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Polyploviricotina Primates Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G43603TQ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2299.86	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G11700UP	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2516.91	Alphavirus Alsuviricetes Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Martellivirales Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Orthornavirae Primates Rattus Riboviria Rodentia Sarcopterygii Semliki Forest virus Simiiformes Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms
G98559XP	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2516.91	Alphavirus Alsuviricetes Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Martellivirales Metazoa Opisthokonta Orthornavirae Primates Riboviria Sarcopterygii Semliki Forest virus Simiiformes Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms
G98485VQ	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1786.65	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G01169EW	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1786.65	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G47925EV	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1932.71	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G65294YU	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1932.71	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Negarnaviricota Neognathae Opisthokonta Orthoparamyxovirinae Orthornavirae Paramyxoviridae Phasianidae Phasianinae Primates Respirovirus muris Respirovirus Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms
G73683LQ	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1932.71	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Monjiviricetes Mononegavirales Negarnaviricota Opisthokonta Orthoparamyxovirinae Orthornavirae Paramyxoviridae Primates Respirovirus muris Respirovirus Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G87690VK	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1934.72	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G53889SN	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2078.77	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G41821MR	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2078.77	Amniota Archelosauria Archosauria Articulavirales Aves Bifurcata Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Elapidae Elapinae Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Lepidosauria Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Naja kaouthia Naja Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthoparamyxovirinae Orthornavirae Paramyxoviridae Phasianidae Phasianinae Polyploviricotina Primates Respirovirus muris Respirovirus Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Theropoda Toxicofera Unidentata Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G68765US	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2080.78	Amniota Articulavirales Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Polyploviricotina Primates Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G38271CW	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	Lewis x Lex N-Glycan core basic N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1567.58	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G24668UI	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan core basic N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1567.58	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Negarnaviricota Opisthokonta Orthoparamyxovirinae Orthornavirae Paramyxoviridae Primates Respirovirus muris Respirovirus Riboviria Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified entries unclassified sequences unidentified
G97529OC	Gal(b1-4)[Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	750.29	Filoviridae Haploviricotina Marburgvirus Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Riboviria Viruses
G88263BY	Gal(b1-4)[GlcNAc(b1-6)]GalNAc-ol		Linkage defined saccharide	588.24	Filoviridae Haploviricotina Marburgvirus Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Riboviria Viruses