GlyCosmos Portal

Source	Last Updated
GlyTouCan	December 15, 2025

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GlyTouCan ID	IUPAC Condensed	Motifs	Archetype	Subsumption Level	Monoisotopic Mass	Organisms
G36157OV	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	G69438ML	Fully-defined saccharide	2093.74	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69533GA	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN		Linkage-defined saccharide	1917.71	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G99129GB	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN		Linkage-defined saccharide	1478.54	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G54884LT	GalNAc(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	LDN LacdiNAc N-Glycan complex		Linkage-defined saccharide	1722.65	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G75151NJ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc	N-acetyllactosamine Type 2 LN		Linkage-defined saccharide	1802.64	Amniota Bacteria Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Pseudomonadota Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G46946XS	Neu5Gc(a2-3)[GalNAc(b1-4)]GalNAc(b1-3)Gal(b1-4)Gal(b1-3)[Neu5Gc(a2-8)Neu5Gc(a2-6)]GalNAc-ol			Linkage-defined saccharide	1874.64	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Teleostei Teleostomi Vertebrata cellular organisms
G03796EF	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-3)GalNAc-ol	Blood group H (type 2) Blood group H (type 2) Leb Lewis b Ley		Linkage-defined saccharide	1042.41	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G14696LD	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[Man(a1-3)[Man(a1-6)]Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	G74022UR	Fully-defined saccharide	1890.66	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10691MJ	Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	G06279KD	Fully-defined saccharide	1989.73	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G27561QX	Gal(?1-4)GlcNAc(?1-2)Man(a1-3)[Gal(?1-4)GlcNAc(?1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(?1-6)]GlcNAc(b1-	N-Glycan core basic		Linkage-defined saccharide	1786.65	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G85079IJ	Neu5Gc(a2-3)Gal(b1-3)[Neu5Gc(a2-6)]GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	G07937GC	Fully-defined saccharide	997.32	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G53868KU	Man(b1-2)Man(b1-2)Man(a1-2)Man(a1-2)Man(a1-	Mannose O-Mannose	G67803PY	Fully-defined saccharide	828.27	Amniota Ascomycota Bilateria Boreoeutheria Candida tropicalis Candida/Lodderomyces clade Candida Catarrhini Chordata Craniata Debaryomycetaceae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Pichiomycetes Primates Saccharomycotina Sarcopterygii Serinales Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G61002BJ		N-Glycan complex N-acetyllactosamine Type 2 LN		Linkage-defined saccharide	2442.88	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Opisthokonta Rodentia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G97607IA	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN		Linkage-defined saccharide	2079.76	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G83006TR	GlcNAc(b1-4)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	G43519BI	Fully-defined saccharide	1478.54	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G15169WU				Monosaccharide composition	3025.07	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G41389XC	Neu5Ac4Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac4Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	G24028ZC	Fully-defined saccharide	2306.8	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Perissodactyla Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G32556JI	Gal(b1-3)GlcNAc(b1-3)[Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)]GalNAc-ol	Sialyl Lewis x Slex Type 1 LN		Linkage-defined saccharide	1390.52	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G42349ED		Lex-Lex		Fully-defined saccharide		Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G49299WO	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Gal(b1-3)Gal(b1-4)Glc(b1-	Lactosylceramide Polysialic acid	G85416ZI	Fully-defined saccharide	1816.62	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G41923EO	Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	G19839OY	Linkage-defined saccharide	1843.67	Batoidea Bilateria Chondrichthyes Chordata Craniata Deuterostomia Elasmobranchii Eukaryota Eumetazoa Gnathostomata Metazoa Opisthokonta Tetronarce californica Tetronarce Torpedinidae Torpediniformes Vertebrata cellular organisms
G87406OV		N-Glycan complex Polysialic acid		Fully-defined saccharide	2296.82	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G37552GO	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[GalNAc(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan complex N-acetyllactosamine Type 2 LN	G31515BE	Linkage-defined saccharide	2087.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10236SD	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	Sialyl Lewis x Slex		Linkage-defined saccharide	3027.08	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G60233QP	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN		Linkage-defined saccharide	2005.72	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms

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