GlyCosmos Portal

Table Filtering

GlyTouCan ID

IUPAC Condensed

Motifs

Subsumption Level

Monoisotopic Mass

Organisms

Taxonomy IDs

Tissues

Display
results per page.

Clear

Submissions

GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1201 - 1225** of **1270** in total

« First

‹ Prev

…

45

46

47

48

49

50

51

Next ›

Last »
GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G71457YN	Neu5Gc(a2-3/6)GalNAc(?1-		Linkage defined saccharide	528.18	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G15807NZ	Neu5Ac(a2-6)GalNAc		Linkage defined saccharide	512.19	Acomys russatus Acomys Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deomyinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G37584UP	Neu5Ac(a2-3/6)GalNAc(?1-		Linkage defined saccharide	512.19	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G08371CD	Neu5Ac(a2-6)GalNAc(?1-		Linkage defined saccharide	512.19	Actinopteri Actinopterygii Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates palustris Lithobates Macaca radiata Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Pecora Primates Protacanthopterygii Ranidae Ranoidea Rattus norvegicus Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G19399OS			Glycosidic topology	512.19	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G36123IU	Neu5Ac(a2-6)GalNAc(a1-	Sialyl Tn antigen	Linkage defined saccharide	512.19	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cestoda Chordata Craniata Cyclophyllidea Deuterostomia Digenea Dipnotetrapodomorpha Discoba Echinostomata Echinostomatoidea Euarchontoglires Eucestoda Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Fasciola hepatica Fasciola Fasciolidae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Kinetoplastea Lagomorpha Laurasiatheria Lentivirus Leporidae Lophotrochozoa Mammalia Mesocestoides vogae Mesocestoides Mesocestoididae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Oryctolagus cuniculus Oryctolagus Pararnavirae Pecora Plagiorchiida Platyhelminthes Primate lentivirus group Primates Protostomia Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Schizotrypanum Simiiformes Spiralia Teleostomi Tetrapoda Theria Trematoda Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G67772KL	Neu5Ac(a2-?)GalNAc(a1-	Sialyl Tn antigen	Linkage defined saccharide	512.19	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00059MO	Gal(a1-4)Gal(b1-4)Glc(b1-	Gb3 Pk antigen globotriaosylceramide	Linkage defined saccharide	504.17	Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Betaproteobacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus sabaeus Chlorocebus Chordata Coelurosauria Craniata Cricetidae Cricetinae Deuterostomia Dinosauria Dipnotetrapodomorpha Discoba Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Glires Gnathostomata Haemophilus influenzae Haemophilus Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kinetoplastea Lagomorpha Laurasiatheria Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Leporidae Macaca fuscata Macaca mulatta Macaca Mammalia Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neisseria gonorrhoeae 1291 Neisseria gonorrhoeae Neisseria Neisseriaceae Neisseriales Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pasteurellaceae Pasteurellales Pecora Phasianidae Phasianinae Primates Pseudomonadota Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Trypanosomatida Trypanosomatidae Vertebrata cellular organisms
G61055FR	Gal(b1-4)Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	504.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G78074HX	Gal(a1-3)Gal(b1-4)Glc(b1-	Lactosylceramide a-Gal antigen	Linkage defined saccharide	504.17	Amniota Artiodactyla Bilateria Boreoeutheria Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G18464VS	GalNAc4S(b1-4)GlcNAc(?1-	4O-sulfated LDN	Linkage defined saccharide	504.13	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G61399NC	Fuc(a1-2)Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	488.17	Amniota Aplysia juliana Aplysia Aplysiida Aplysiidae Aplysioidea Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Euthyneura Gastropoda Glires Gnathostomata Heterobranchia Laurasiatheria Lophotrochozoa Mammalia Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Protostomia Rattus Rodentia Sarcopterygii Spiralia Suidae Suina Sus scrofa Sus Tectipleura Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G39807DF	Neu5Gc(a2-6)Gal(?1-		Linkage defined saccharide	487.15	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G77824XV	Neu5Ac(a2-3)Gal(?1-		Linkage defined saccharide	471.16	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G35815VK	Neu5Ac(a2-?)Gal(b1-	GM4	Linkage defined saccharide	471.16	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G30207PZ	Neu5Ac(a2-3)Gal(b1-	GM4	Linkage defined saccharide	471.16	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cetacea Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Zeiogadaria cellular organisms
G44345UH	Neu5Ac(a2-6)Gal(?1-		Linkage defined saccharide	471.16	Amniota Amphiesmenoptera Amphipyrinae Arthropoda Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Pecora Primates Protostomia Pterygota Rodentia Ruminantia Sarcopterygii Simiiformes Spodoptera frugiperda Spodoptera Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G71814NS	Neu5Ac(a2-3)Gal(a1-		Linkage defined saccharide	471.16	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G75529YY	Neu5Ac(a2-6)Gal		Linkage defined saccharide	471.16	Acomys russatus Acomys Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deomyinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G45388GV	GalNAc(a1-3)GalNAc(?1-	O-Glycan core 5 fuzzy	Linkage defined saccharide	424.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gerbillinae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Meriones unguiculatus Meriones Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G71838YU			Glycosidic topology	424.17	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G91311MD	GalNAc(a1-3)GlcNAc		Linkage defined saccharide	424.17	Acomys russatus Acomys Amniota Bilateria Boreoeutheria Chordata Craniata Deomyinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G97304IA	GlcNAc(b1-4)GlcNAc		Linkage defined saccharide	424.17	Acomys russatus Acomys Amniota Bacteria Betaproteobacteria Bilateria Boreoeutheria Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Craniata Deomyinae Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Epsilonproteobacteria Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gammaproteobacteria Glires Gnathostomata Haemophilus influenzae Haemophilus Haplorrhini Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neisseria gonorrhoeae Neisseria Neisseriaceae Neisseriales Opisthokonta Pasteurellaceae Pasteurellales Primates Pseudomonadota Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G77259GO	Gal(?1-?)GalNAc-ol		Glycosidic topology	385.16	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G67200RM	Gal(?1-?)GalNAc(?1-		Glycosidic topology	383.14	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms