GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	February 17, 2025

Displaying entries **101 - 125** of **168** in total

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GlyTouCan ID	IUPAC Condensed	Motifs ▲	Subsumption Level	Monoisotopic Mass	Organisms
G54831HN	GlcNAc(b1-2/4)Gal(b1-4)GlcNAc(b1-3/6)[Gal(b1-4)GlcNAc(a1-3)Gal(b1-3/6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	1318.5	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G67208MP	Fuc(a1-2)Gal(b1-4)GlcNAc(a1-3)Gal(b1-3/6)[GlcNAc(a1-2/4)Gal(b1-4)GlcNAc(b1-3/6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	1464.56	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70520NA	GlcNAc(a1-3/4)Gal(b1-3/4)Gal(b1-3)[GlcNAc(a1-4)Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	1318.5	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84273UX	GlcNAc(b1-3/4)Gal(b1-3/4)Gal(b1-3)[GlcNAc(b1-4)Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	1318.5	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G58793GF	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GalNAc(?1-	O-Glycan core 1 fuzzy Polysialic acid	Linkage-defined saccharide	1039.37	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G67326LW	Neu5Gc(a2-3)Gal(b1-3)[Neu5Gc(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage-defined saccharide	997.32	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Perissodactyla Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G91484JV	Neu5Gc(a2-3)Gal(b1-3)GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage-defined saccharide	690.23	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Perissodactyla Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G22412KQ	GlcNAc(b1-2/3/4)Gal(b1-3/6)[GlcNAc(b1-3/6)]GalNAc(?1-	O-Glycan core 2 fuzzy O-Glycan core 3 fuzzy	Linkage-defined saccharide	789.3	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G82950HY	Gal(b1-3)[GlcNAc(b1-6)]GalNAc(?1-	O-Glycan core 2 fuzzy	Linkage-defined saccharide	586.22	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates palustris Lithobates sphenocephalus Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Opisthokonta Ovis aries Ovis Pecora Pipidae Pipoidea Primates Rana arvalis Rana Rana Ranidae Ranoidea Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus artificial sequences cellular organisms other entries other sequences synthetic construct
G00034MO	Gal(b1-3)[GlcNAc(b1-6)]GalNAc	O-Glycan core 2 fuzzy	Linkage-defined saccharide	586.22	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ovis aries Ovis Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G30425ZJ	GlcNAc(b1-3/4)Gal(b1-3)[GlcNAc(b1-6)]GalNAc(?1-	O-Glycan core 2 fuzzy	Linkage-defined saccharide	789.3	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G85714GY	GlcNAc(b1-3)GalNAc(?1-	O-Glycan core 3 fuzzy	Linkage-defined saccharide	424.17	Ambystoma mexicanum Ambystoma Ambystomatidae Amniota Amphibia Artiodactyla Ascidiacea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Caudata Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ovis aries Ovis Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Salamandroidea Sarcopterygii Simiiformes Stolidobranchia Styela plicata Styela Styelidae Teleostomi Tetrapoda Theria Tunicata Vertebrata cellular organisms
G18892PE	GlcNAc(b1-3)[GlcNAc(b1-6)]GalNAc(?1-	O-Glycan core 4 fuzzy	Linkage-defined saccharide	627.25	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ovis aries Ovis Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59126YU	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-	O-Linked mannose	Fully-defined saccharide	836.29	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01767JJ	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside Polysialic acid	Fully-defined saccharide	998.34	Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Betaproteobacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Macaca fuscata Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neisseria meningitidis Neisseria Neisseriaceae Neisseriales Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Primates Pseudomonadota Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G86143FV	Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside Type 1 LN	Fully-defined saccharide	1072.38	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G60645OK	Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside a-Gal antigen	Fully-defined saccharide	869.3	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carnivora Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Phasianidae Phasianinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G58896AZ	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Fully-defined saccharide	707.25	Amniota Archelosauria Archosauria Artiodactyla Aves Bacilli Bacillota Bacteria Betaproteobacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carnivora Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lactobacillales Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neisseria gonorrhoeae Neisseria Neisseriaceae Neisseriales Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Pseudomonadota Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Streptococcaceae Streptococcus pneumoniae Streptococcus Suidae Suina Sus scrofa Sus Teleostomi Terrabacteria group Tetrapoda Theria Theropoda Vertebrata cellular organisms
G69161BS	Neu5Gc(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Fully-defined saccharide	1321.43	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Carnivora Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G81009UW	Neu5Ac(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Fully-defined saccharide	1305.43	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Carnivora Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86874HJ	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	Polysialic acid	Linkage-defined saccharide	2224.8	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G57321FI	GalNAc(a1-	Tn antigen			Acalyptratae Amniota Armadillidae Armadillo Arthropoda Artiodactyla Artverviricota Ascaridoidea Ascaridomorpha Ascomycota Bacteria Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenorhabditis elegans Caenorhabditis Calliphoridae Calyptratae Caprinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cestoda Chlorocebus aethiops Chlorocebus Chordata Chromadorea Chrysomyinae Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crinocheta Crustacea Cyclophyllidea Cyclorrhapha Deuterostomia Dicondylia Digenea Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Echinococcus granulosus group Echinococcus granulosus Echinococcus Echinostomata Echinostomatoidea Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eucestoda Eukaryota Eumalacostraca Eumetazoa Euteleostomi Eutheria Fasciola hepatica Fasciola Fasciolidae Fungi Gammaproteobacteria Glires Gnathostomata Haplorrhini Heligmosomidae Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Insecta Isopoda Lagomorpha Laurasiatheria Lentivirus Leporidae Lophotrochozoa Macaca mulatta Macaca Malacostraca Mammalia Mandibulata Mesocestoides corti Mesocestoides vogae Mesocestoides Mesocestoididae Metazoa Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Nematoda Neoptera Nidovirales Nippostrongylus brasiliensis Nippostrongylus Oestroidea Oniscidea Opisthokonta Ortervirales Orthocoronavirinae Orthoretrovirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Panarthropoda Pancrustacea Pararnavirae Pecora Peloderinae Peracarida Phormini Pisoniviricetes Pisuviricota Plagiorchiida Platyhelminthes Primate lentivirus group Primates Protophormia terraenovae Protophormia Protostomia Pseudomonadaceae Pseudomonadales Pseudomonadota Pseudomonas aeruginosa group Pseudomonas aeruginosa Pseudomonas Pterygota Rattus norvegicus Rattus Retroviridae Revtraviricetes Rhabditida Rhabditidae Rhabditina Rhabditoidea Rhabditomorpha Riboviria Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Spiralia Spirurina Strongyloidea Taenia hydatigena Taenia Taeniidae Teleostomi Tetrapoda Theria Toxocara canis Toxocara Toxocaridae Trematoda Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup saccharomyceta unclassified Primate lentivirus group
G23799PV	Gal(b1-3)GlcNAc(b1-6)Gal(b1-4)[Fuc(a1-3)]Glc	Type 1 LN	Linkage-defined saccharide	853.31	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Capra hircus Capra Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G77070UX	Gal(a1-3)Gal(b1-4)Glc	a-Gal antigen	Linkage-defined saccharide	504.17	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Capra hircus Capra Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01347XI	Gal(a1-4)Gal(b1-4)GlcNAc		Linkage-defined saccharide	545.2	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms