GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1776 - 1800** of **2251** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G48856VC	HexNAc(b1-?)Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	951.35	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G80193TN	Gal3S(b1-3)GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc(b1-	Gb4 Globoside P antigen	Linkage defined saccharide	949.26	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G97310HO	Gal3S(b1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)Glc(b1-	Cytolipin R Mono-sulfated globopentaosylceramide	Linkage defined saccharide	949.26	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G27243GN	Neu5Ac(a2-8)Neu5Gc(a2-3)Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	940.3	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Carnivora Catarrhini Cetacea Chordata Craniata Delphinidae Delphinus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Odontoceti Opisthokonta Ovis aries Ovis Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Stenella coeruleoalba Stenella Teleostomi Tetrapoda Theria Vertebrata Whippomorpha cellular organisms
G80829RZ	Neu5Gc(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	GM3	Linkage defined saccharide	940.3	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G80577JZ	GlcNAc(b1-3)[Fuc(a1-2)Gal(b1-?)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	937.38	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G76874HM	Fuc(a1-2)Gal(b1-?)GlcNAc(b1-3)[GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	937.38	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00018MO	GlcA(b1-3)GalNAc4S(b1-4)GlcA(b1-3)GalNAc4S		Linkage defined saccharide	936.15	Amniota Arthropoda Bilateria Boreoeutheria Brachyura Chaceon affinis Chaceon Chondrichthyes Chordata Craniata Crustacea Decapoda Deuterostomia Dipnotetrapodomorpha Ecdysozoa Elasmobranchii Euarchontoglires Eubrachyura Eucarida Eukaryota Eumalacostraca Eumetazoa Euteleostomi Eutheria Geryonidae Glires Gnathostomata Heterotremata Malacostraca Mammalia Mandibulata Metazoa Multicrustacea Muridae Murinae Muroidea Myomorpha Opisthokonta Panarthropoda Pancrustacea Pleocyemata Portunoidea Protostomia Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G31061SB	GlcNAc(b1-3)[Fuc(a1-2)Gal(b1-?)GlcNAc(b1-6)]GalNAc(?1-	N-acetyllactosamine O-Glycan core 4 fuzzy Type 1 LN Type 2 LN	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04908IA	GalNAc(?1-3)[Fuc(?1-2)]Gal(?1-3)GlcNAc(?1-3)GalNAc(a1-	Tn antigen	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23990GP	Fuc(?1-2)Gal(?1-?)GlcNAc(?1-3)[GlcNAc(?1-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68931EB	Fuc(a1-2)Gal(b1-?)GlcNAc(b1-3)[GlcNAc(b1-6)]GalNAc(?1-	N-acetyllactosamine O-Glycan core 4 fuzzy Type 1 LN Type 2 LN	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G82698QQ	GalNAc(?1-3)[Fuc(?1-2)]Gal(?1-3)[GlcNAc(?1-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G96416NB	GlcNAc(?1-3)[Fuc(?1-2)Gal(?1-?)GlcNAc(?1-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	935.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G98544DH	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	9-O-Acetyl GD3 GD3	Linkage defined saccharide	924.31	Actinopteri Actinopterygii Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Balaenoptera edeni Balaenoptera Balaenopteridae Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chinchilla Chinchillidae Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leucoraja erinacea Leucoraja Macaca fuscata Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Narcinidae Neognathae Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Papio hamadryas Papio Pecora Phasianidae Phasianinae Pipidae Pipoidea Primates Rajidae Rajiformes Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Vertebrata Whippomorpha Xenopodinae Xenopus cellular organisms
G83960NP	Neu5Ac(a2-?)[Neu5Ac(a2-?)Gal(b1-3)]Gal(?1-	GM4	Linkage defined saccharide	924.31	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G62167DK	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	912.34	Aculeata Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Anthophila Apidae Apocrita Apoidea Archelosauria Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Primates Protostomia Pterygota Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G64239KZ	Gal(b1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	912.34	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G07430HH	Gal(b1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc(?1-	N-acetyllactosamine O-Glycan core 2 fuzzy Type 2 LN a-Gal antigen	Linkage defined saccharide	910.33	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G14669DU			Glycosidic topology	910.33	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G15301IS	GalNAc(a1-3)GalNAc(b1-4)Gal(a1-4)Gal(b1-4)Glc(b1-	Gb3 Pk antigen globotriaosylceramide	Linkage defined saccharide	910.33	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G20624LQ	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	910.33	50 kb inversion clade Aculeata Agelenidae Agelenopsis aperta Agelenopsis Amniota Amphibia Amphiesmenoptera Amphipyrinae Anthophila Anura Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arachnida Araneae Araneomorphae Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chelicerata Chordata Coelurosauria Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Entelegynae Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fabaceae Fabales Fabeae Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Heteroneura Hexapoda Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera IRL clade Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Lophotrochozoa Magnoliopsida Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Panpulmonata Papilionoideae Pecora Pentapetalae Perdicinae Phaseoleae Phaseolus Phasianidae Phasianinae Pisum sativum Pisum Plusiinae Primates Protostomia Pterygota RTA clade Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spiralia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Stylommatophora Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids rosids unclassified entries unclassified sequences unidentified
G22768VO	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	910.33	50 kb inversion clade Acalyptratae Amniota Amphibia Anura Arthropoda Artiodactyla Batrachia Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Neobatrachia Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rana Ranidae Ranoidea Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids
G25323VU	Gal(b1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN a-Gal antigen	Linkage defined saccharide	910.33	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G25413GH			Linkage defined saccharide	910.33	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms