GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1 - 25** of **235** in total

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GlyTouCan ID	IUPAC Condensed ▲	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G17475NE	All(b1-				Arthropoda Bilateria Chrysolina fuliginosa Chrysolina Chrysomelidae Chrysomelinae Chrysomelini Chrysomeloidea Coleoptera Cucujiformia Dicondylia Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Insecta Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Polyphaga Protostomia Pterygota cellular organisms
G39785TO	Gal(a1-3)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	910.33	Arthropoda Ascarididae Ascaridoidea Ascaridomorpha Ascaris suum Ascaris Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Chromadorea Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Nematoda Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Rhabditida Schizophora Spirurina cellular organisms
G94613NP	Gal(a1-4)GalNAc(a1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	1113.41	Arthropoda Bilateria Brachycera Calliphoridae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G78436BB	Gal(b1-3)GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	383.14	Amniota Amphiesmenoptera Amphipyrinae Arthropoda Artiodactyla Bilateria Boreoeutheria Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gerbillinae Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Insecta Lagomorpha Laurasiatheria Lepidoptera Leporidae Mammalia Mandibulata Meriones unguiculatus Meriones Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Oryctolagus cuniculus Oryctolagus Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus Rodentia Sarcopterygii Simiiformes Spodoptera frugiperda Spodoptera Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00031MO	Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	383.14	Acalyptratae Acanthomorphata Actinopteri Actinopterygii Amniota Arthropoda Artiodactyla Bacteria Bacteroidaceae Bacteroidales Bacteroides ovatus Bacteroides Bacteroidia Bacteroidota/Chlorobiota group Bacteroidota Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria FCB group Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Neopterygii Neoteleostei Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Panarthropoda Pancrustacea Paracanthopterygii Pecora Pisoniviricetes Pisuviricota Pollachius pollachius Pollachius Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Shigella dysenteriae Shigella Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses Zeiogadaria cellular organisms melanogaster group melanogaster subgroup
G28484YT	Gal(b1-3)GalNAc(a1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	1113.41	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G63328VH	Gal(b1-3)GalNAc(a1-4)GalNAc(b1-4)GlcNAc6PEtn(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	1236.42	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G01534TU	Gal(b1-3)GalNAc(b1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	383.14	Amniota Amphiesmenoptera Amphipyrinae Arthropoda Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lepidoptera Mammalia Mandibulata Metazoa Muroidea Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rodentia Sarcopterygii Simiiformes Spodoptera frugiperda Spodoptera Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G55908IH	Gal(b1-3)GalNAc(b1-3)GlcNAc(b1-3)Gal(b1-3)GalNAc(a1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	1681.62	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G83691NP	Gal(b1-3)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	910.33	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G51191BM	Gal(b1-3)GalNAc(b1-4)GlcNAc6PEtn(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	1033.34	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G00033MO	Gal(b1-3)[GlcNAc(b1-6)]GalNAc(a1-	O-Glycan core 2	Linkage defined saccharide	586.22	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G79333MY	Gal(b1-4)GlcNAc(a1-		Linkage defined saccharide	383.14	Amniota Armadillidae Armadillo Arthropoda Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crinocheta Crustacea Deuterostomia Dipnotetrapodomorpha Discoba Ecdysozoa Euarchontoglires Euglenozoa Eukaryota Eumalacostraca Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Isopoda Kinetoplastea Laurasiatheria Malacostraca Mammalia Mandibulata Metakinetoplastina Metazoa Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Oniscidea Opisthokonta Panarthropoda Pancrustacea Peracarida Primates Protostomia Rattus norvegicus Rattus Rodentia Sarcopterygii Schizotrypanum Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata cellular organisms
G57321FI	GalNAc(a1-	Tn antigen			Acalyptratae Amniota Armadillidae Armadillo Arthropoda Artiodactyla Artverviricota Ascaridoidea Ascaridomorpha Ascomycota Bacteria Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenorhabditis elegans Caenorhabditis Calliphoridae Calyptratae Caprinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cestoda Chlorocebus aethiops Chlorocebus Chordata Chromadorea Chrysomyinae Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crinocheta Crustacea Cyclophyllidea Cyclorrhapha Deuterostomia Dicondylia Digenea Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Echinococcus granulosus group Echinococcus granulosus Echinococcus Echinostomata Echinostomatoidea Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eucestoda Eukaryota Eumalacostraca Eumetazoa Euteleostomi Eutheria Fasciola hepatica Fasciola Fasciolidae Fungi Gammaproteobacteria Glires Gnathostomata Haplorrhini Heligmosomidae Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Insecta Isopoda Lagomorpha Laurasiatheria Lentivirus Leporidae Lophotrochozoa Macaca mulatta Macaca Malacostraca Mammalia Mandibulata Mesocestoides corti Mesocestoides vogae Mesocestoides Mesocestoididae Metazoa Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Nematoda Neoptera Nidovirales Nippostrongylus brasiliensis Nippostrongylus Oestroidea Oniscidea Opisthokonta Ortervirales Orthocoronavirinae Orthoretrovirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Panarthropoda Pancrustacea Pararnavirae Pecora Peloderinae Peracarida Phormini Pisoniviricetes Pisuviricota Plagiorchiida Platyhelminthes Primate lentivirus group Primates Protophormia terraenovae Protophormia Protostomia Pseudomonadaceae Pseudomonadales Pseudomonadota Pseudomonas aeruginosa group Pseudomonas aeruginosa Pseudomonas Pterygota Rattus norvegicus Rattus Retroviridae Revtraviricetes Rhabditida Rhabditidae Rhabditina Rhabditoidea Rhabditomorpha Riboviria Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Spiralia Spirurina Strongyloidea Taenia hydatigena Taenia Taeniidae Teleostomi Tetrapoda Theria Toxocara canis Toxocara Toxocaridae Trematoda Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup saccharomyceta unclassified Primate lentivirus group
G30781KG	GalNAc(a1-4)GalNAc(b1-		Linkage defined saccharide	424.17	Arthropoda Bilateria Camptochironomus Chironomidae Chironominae Chironomini Chironomoidea Chironomus tentans Chironomus Culicomorpha Dicondylia Diptera Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Insecta Mandibulata Metazoa Nematocera Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota cellular organisms
G66868KB	GalNAc(a1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	951.35	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G94095NQ	GalNAc(a1-4)GalNAc(b1-4)GlcNAc6PEtn(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	1074.36	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G03732ES	GalNAc(b1-3)GlcNAc(b1-3)Gal(b1-3)GalNAc(a1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	1519.57	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G50813ST	GalNAc(b1-4)GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	951.35	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G40793GY	GalNAc(b1-4)GlcNAc(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide LDN LacdiNAc	Linkage defined saccharide	748.27	Arthropoda Bilateria Bivalvia Brachycera Calliphoridae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lophotrochozoa Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Mollusca Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora Spiralia cellular organisms
G30055NL	GalNAc(b1-4)GlcNAc6PEtn(b1-3)Man(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	871.28	Arthropoda Bilateria Brachycera Calliphora vicina Calliphora Calliphoridae Calliphorinae Calyptratae Cyclorrhapha Dicondylia Diptera Ecdysozoa Endopterygota Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Lucilia caesar Lucilia sericata Lucilia Luciliinae Mandibulata Metazoa Muscomorpha Neoptera Oestroidea Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora cellular organisms
G02240AO	GalNAc(b1-4)[Fuc(a1-3)]GlcNAc(?1-	Fucosylated LDN LDNF	Linkage defined saccharide	570.23	Aculeata Amniota Anthophila Apidae Apinae Apini Apis Apocrita Apoidea Arthropoda Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Mammalia Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G80019MD	GalNAc(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	Fucosylated LDN LDNF N-Glycan core basic	Linkage defined saccharide	1462.54	Aculeata Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Bilateria Dicondylia Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Hymenoptera Insecta Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota cellular organisms
G87773EB	GalNAc(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Fucosylated LDN LDNF N-Glycan core basic	Linkage defined saccharide	1462.54	Aculeata Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Bilateria Dicondylia Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Hymenoptera Insecta Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota cellular organisms
G48902RK	GalNAc(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	Fucosylated LDN LDNF	Linkage defined saccharide	1464.56	Aculeata Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Bilateria Dicondylia Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Hymenoptera Insecta Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota cellular organisms