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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1 - 25** of 71 in total

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GlyTouCan ID	IUPAC Condensed ▼	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G61628UG	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	1190.44	Ananas comosus Ananas Bilateria Bromeliaceae Bromelioideae Embryophyta Eukaryota Eumetazoa Euphyllophyta Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Liliopsida Lophotrochozoa Magnoliopsida Mesangiospermae Metazoa Mollusca Opisthokonta Panpulmonata Petrosaviidae Poales Protostomia Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Tracheophyta Viridiplantae cellular organisms commelinids
G13181ZZ	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1188.43	Amniota Apiaceae Apiales Apiineae Apioideae Bilateria Boreoeutheria Catarrhini Chordata Craniata Daucinae Daucus carota Daucus sect. Daucus Daucus Deuterostomia Digenea Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fissurellidae Fissurelloidea Gastropoda Gnathostomata Gunneridae Haplorrhini Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Hominidae Homininae Hominoidea Homo sapiens Homo Lepetellida Lophotrochozoa Magnoliopsida Mammalia Megathura crenulata Megathura Mesangiospermae Metazoa Mollusca Opisthokonta Panpulmonata Pentapetalae Platyhelminthes Primates Protostomia Sarcopterygii Scandiceae Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Simiiformes Spermatophyta Spiralia Streptophyta Streptophytina Strigeidida Stylommatophora Teleostomi Tetrapoda Theria Tracheophyta Trematoda Vertebrata Vetigastropoda Viridiplantae asterids campanulids cellular organisms eudicotyledons
G14307KB	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1188.43	Acalypheae Acalyphoideae Ananas comosus Ananas Apiaceae Apiales Apiineae Apioideae Bilateria Bromeliaceae Bromelioideae Caryophyllaceae Caryophyllales Daucinae Daucus carota Daucus sect. Daucus Daucus Digenea Embryophyta Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euthyneura Gastropoda Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Liliopsida Lophotrochozoa Magnoliopsida Malpighiales Mesangiospermae Metazoa Mollusca Opisthokonta Panpulmonata Pentapetalae Petrosaviidae Platyhelminthes Poales Protostomia Ricinus communis Ricinus Scandiceae Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Silene latifolia subsp. alba Silene latifolia Silene sect. Melandrium Silene subgen. Behenantha Silene Sileneae Spermatophyta Spiralia Streptophyta Streptophytina Strigeidida Stylommatophora Tracheophyta Trematoda Viridiplantae asterids campanulids cellular organisms commelinids eudicotyledons fabids rosids
G75311GD	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	1188.43	50 kb inversion clade Acalypheae Acalyphoideae Bilateria Caryophyllaceae Caryophyllales Embryophyta Erythrina corallodendron Erythrina Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euthyneura Fabaceae Fabales Gastropoda Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Magnoliopsida Malpighiales Mesangiospermae Metazoa Mollusca NPAAA clade Opisthokonta Panpulmonata Papilionoideae Pentapetalae Phaseoleae Protostomia Ricinus communis Ricinus Silene latifolia subsp. alba Silene latifolia Silene sect. Melandrium Silene subgen. Behenantha Silene Sileneae Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G69917FQ	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1044.39	50 kb inversion clade Acalypheae Acalyphoideae Acer pseudoplatanus Acer Acereae Ananas comosus Ananas Bilateria Bromeliaceae Bromelioideae Caragana arborescens Caragana Caraganeae Cercideae Cercidinae Cercidoideae Cladrastis clade Embryophyta Erythrina caffra Erythrina corallodendron Erythrina crista-galli Erythrina latissima Erythrina Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euthyneura Fabaceae Fabales Fabeae Gastropoda Griffonia simplicifolia Griffonia Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Hippocastanoideae Hologalegina IRL clade Lens culinaris Lens Liliopsida Lophotrochozoa Magnoliopsida Malpighiales Mesangiospermae Metazoa Mollusca NPAAA clade Opisthokonta Panpulmonata Papilionoideae Pentapetalae Petrosaviidae Phaseoleae Phaseolus coccineus Phaseolus lunatus Phaseolus vulgaris Phaseolus Platanaceae Platanus Poales Proteales Protostomia Ricinus communis Ricinus Sapindaceae Sapindales Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Styphnolobium japonicum Styphnolobium Tracheophyta Vicia faba Vicia Vigna cylindrica Vigna Viridiplantae Wisteria floribunda Wisteria Wisterieae cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade malvids rosids
G64979EW	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1042.37	50 kb inversion clade Acalypheae Acalyphoideae Amniota BOP clade Bilateria Boreoeutheria Chordata Craniata Cucurbitaceae Cucurbitales Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fabaceae Fabales Gastropoda Glires Gnathostomata Gunneridae Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Hygrophila Liliopsida Lophotrochozoa Lymnaea stagnalis Lymnaea Lymnaeidae Lymnaeoidea Magnoliopsida Malpighiales Mammalia Mesangiospermae Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panpulmonata Papilionoideae Pentapetalae Petrosaviidae Phaseoleae Phaseolus vulgaris Phaseolus Poaceae Poales Protostomia Ricinus communis Ricinus Rodentia Sarcopterygii Solanaceae Solanales Spermatophyta Spiralia Streptophyta Streptophytina Stylommatophora Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae asterids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids rosids
G20624LQ	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	910.33	50 kb inversion clade Aculeata Agelenidae Agelenopsis aperta Agelenopsis Amniota Amphibia Amphiesmenoptera Amphipyrinae Anthophila Anura Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arachnida Araneae Araneomorphae Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chelicerata Chordata Coelurosauria Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Entelegynae Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fabaceae Fabales Fabeae Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Heteroneura Hexapoda Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera IRL clade Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Lophotrochozoa Magnoliopsida Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Panpulmonata Papilionoideae Pecora Pentapetalae Perdicinae Phaseoleae Phaseolus Phasianidae Phasianinae Pisum sativum Pisum Plusiinae Primates Protostomia Pterygota RTA clade Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spiralia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Stylommatophora Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids rosids unclassified entries unclassified sequences unidentified
G07016SY	Man(a1-3)[Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	1948.74	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G35731IU	Man(a1-3)[Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	1946.72	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G84794IE	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2298.86	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G01333FO	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2300.88	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G37009XT	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G44088KZ	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2124.81	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G82902UU	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G43499RV	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G35271HQ	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2298.86	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G02641SK	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2300.88	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G64237KZ	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2298.86	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G96482FU	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2298.86	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G02111QR	Gal4Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1800.67	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G02254OB	Gal4Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	2881.09	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G24712KO	Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	1802.68	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G97467MX	Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2883.1	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G82246KY	Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	2881.09	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G87907JO	Gal3Me(b1-6)Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2152.8	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms