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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1 - 25** of **642** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G23843KC	Man(b1-4)Man(b1-4)[Gal(a1-6)]Man(b1-4)[Gal(a1-6)]Man(b1-4)Man(b1-4)Man(b1-4)[Man(b1-6)]Man(b1-4)[Man(b1-6)]Man(b1-4)Man(b1-4)Man(b1-4)[Gal(a1-6)]Man(b1-4)[Gal(a1-3)Gal(a1-3)Gal(a1-6)]Man(b1-4)Man(b1-4)Man(b1-4)[Man(b1-6)]Man(b1-4)[Man(b1-6)]Man(b1-4)Man(b1-4)Man(b1-4)[Man(b1-6)]Man(b1-4)[Man(b1-6)]Man(b1-4)Man(b1-	Mannose O-Mannose	Linkage defined saccharide	5365.75	Caesalpinioideae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae Mimoseae Pentapetalae Prosopis cineraria Prosopis Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids mimosoid clade rosids
G68578MR	Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	3049.03	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G45660MA	Xyl(a1-6)Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	3033.03	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G68652LV	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	3033.03	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G59478GT	Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	3019.02	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G68258NR	Araf(a1-2)[Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	3019.02	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G16021ZZ	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Araf(a1-3)Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2989.01	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G21498CL	Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2886.98	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G67899KW	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2886.98	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G93638BI	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	50 kb inversion clade Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Butea monosperma Butea Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Papilionoideae Pecora Pentapetalae Phaseoleae Phasianidae Phasianinae Primates Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids unclassified entries unclassified sequences unidentified
G08732ZV	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2856.97	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G41817FI	Araf(a1-2)[Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2856.97	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G04911WU	Xyl(a1-6)Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2724.92	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G25780MO	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2710.91	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G28534IJ	Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2710.91	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G54753GT	Araf(a1-2)[Xyl(a1-6)Glc(b1-4)[Gal(b1-2)Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2710.91	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G51394RI	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Araf(a1-3)Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2680.9	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G28705ZJ	Araf(a1-2)[Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2548.86	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G29337EE	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2548.86	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G30881UU	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2548.86	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G74541SE	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc(b1-4)[Xyl(b1-2)][Xyl(a1-6)]Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)[Fuc(a1-2)Gal(b1-2)Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	2548.86	Acer pseudoplatanus Acer Acereae Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G21656PI	Gal(?1-3)[Gal(?1-4)]GlcNAc(?1-4)[Fuc(a1-3)]GlcNAc(?1-3)Man(a1-3)[Gal(?1-3)Man(a1-3)Man(a1-6)][Xyl(?1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan core basic	Linkage defined saccharide	2388.86	50 kb inversion clade Arachis hypogaea Arachis Dalbergieae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae Papilionoideae Pentapetalae Pterocarpus clade Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms dalbergioids sensu lato eudicotyledons fabids rosids
G09568JT	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	50 kb inversion clade Amniota Batoidea Bilateria Boreoeutheria Butea monosperma Butea Catarrhini Chondrichthyes Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elasmobranchii Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Magnoliopsida Mammalia Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Primates Rodentia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G72954XC	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2368.84	50 kb inversion clade Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Butea monosperma Butea Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neognathae Opisthokonta Papilionoideae Pecora Pentapetalae Perissodactyla Phaseoleae Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids unclassified entries unclassified sequences unidentified
G88958OW	Gal(b1-4)[Fuc(a1-6)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-6)]GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2212.82	Acer pseudoplatanus Acer Acereae Acrogymnospermae Conifers II Cryptomeria japonica Cryptomeria Cupressaceae Cupressales Embryophyta Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Pinidae Pinopsida Platanaceae Platanus Proteales Sapindaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids