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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 4, 2025

Displaying **all 17** entries
GlyTouCan ID	IUPAC Condensed	Motifs	Archetype	Subsumption Level	Monoisotopic Mass	Organisms ▲
G20624LQ	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	G00026MO	Linkage-defined saccharide	910.33	50 kb inversion clade Aculeata Agelenidae Agelenopsis aperta Agelenopsis Amniota Amphibia Amphiesmenoptera Amphipyrinae Anthophila Anura Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arachnida Araneae Araneomorphae Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chelicerata Chordata Coelurosauria Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Entelegynae Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eupulmonata Euteleostomi Eutheria Euthyneura Fabaceae Fabales Fabeae Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Heteroneura Hexapoda Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera IRL clade Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Lophotrochozoa Magnoliopsida Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Panpulmonata Papilionoideae Pecora Pentapetalae Perdicinae Phaseoleae Phaseolus Phasianidae Phasianinae Pisum sativum Pisum Plusiinae Primates Protostomia Pterygota RTA clade Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spiralia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Stylommatophora Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Viridiplantae asterids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids rosids unclassified entries unclassified sequences unidentified
G70073SG	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	G42680EX	Linkage-defined saccharide	1113.41	Acer pseudoplatanus Acer Acereae Achelata Alpharhabdovirinae Amniota Amphibia Amphiesmenoptera Amphipyrinae Anas sp. Anas Anatidae Anatinae Anseriformes Anura Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crustacea Decapoda Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Euarchontoglires Eucarida Eukaryota Eumalacostraca Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Magnoliopsida Malacostraca Mammalia Mandibulata Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Orthornavirae Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pecora Pentapetalae Perdicinae Phasianidae Phasianinae Pleocyemata Plusiinae Primates Protostomia Pterygota Ranidae Ranoidea Rhabdoviridae Riboviria Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids cellular organisms eudicotyledons lamiids malvids rosids unclassified Anas unclassified Vesiculovirus
G38297TU	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	G78341DP	Linkage-defined saccharide	1437.51	Acer pseudoplatanus Acer Acereae Actinopteri Actinopterygii Alpharhabdovirinae Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicales Carica papaya Carica Caricaceae Catarrhini Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Haploviricotina Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Magnoliopsida Mammalia Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neobatrachia Neognathae Neopterygii Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Orthornavirae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Pentapetalae Phasianidae Phasianinae Primates Ranidae Ranoidea Rhabdoviridae Riboviria Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids cellular organisms eudicotyledons lamiids malvids rosids unclassified Vesiculovirus
G04445KN	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	G37841NF	Linkage-defined saccharide	1275.46	Acer pseudoplatanus Acer Acereae Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicales Carica papaya Carica Caricaceae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Gunneridae Haplorrhini Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Magnoliopsida Mammalia Mesangiospermae Metazoa Neobatrachia Neognathae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Opisthokonta Pecora Pentapetalae Phasianidae Phasianinae Primates Ranidae Ranoidea Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae asterids cellular organisms eudicotyledons lamiids malvids rosids
G44747CU	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	G67924FL	Linkage-defined saccharide	1599.57	Actinopteri Actinopterygii Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Artverviricota Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Coelurosauria Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Laurasiatheria Lentivirus Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neognathae Neopterygii Opisthokonta Ortervirales Orthoretrovirinae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pararnavirae Pecora Phasianidae Phasianinae Primate lentivirus group Primates Ranidae Ranoidea Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G86386QT	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	G00978TF	Linkage-defined saccharide	1275.46	Amniota Amphibia Anas platyrhynchos Anas sp. Anas Anatidae Anatinae Anseriformes Anura Archelosauria Archosauria Aves Batrachia Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Leporidae Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Perdicinae Phasianidae Phasianinae Primates Ranidae Ranoidea Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified Anas unclassified entries unclassified sequences unidentified
G61751GZ	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	G42680EX	Fully-defined saccharide	1113.41	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G69411IG	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	G00978TF	Fully-defined saccharide	1275.46	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G08520NM	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	G78341DP	Fully-defined saccharide	1437.51	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms
G56756FT	Man(a1-3)[Man(a1-6)]Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		G76938LQ	Linkage-defined saccharide	1072.38	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bovidae Caprinae Carnivora Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muroidea Myomorpha Neobatrachia Neognathae Opisthokonta Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G16828VN	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	G67924FL	Fully-defined saccharide	1599.57	Amniota Amphibia Anura Archelosauria Archosauria Artverviricota Aves Batrachia Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Lentivirus Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neognathae Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Phasianidae Phasianinae Primate lentivirus group Primates Rana Ranidae Ranoidea Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G94106MV	Man(a1-3)[Man(a1-6)]Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-		G76938LQ	Fully-defined saccharide	1072.38	Amniota Amphibia Anura Archelosauria Archosauria Aves Batrachia Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G47498FQ	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	G04851FX	Linkage-defined saccharide	1437.51	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Lithobates pipiens Lithobates Mammalia Metazoa Neobatrachia Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Ranidae Ranoidea Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G95956WE	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	G17226QK	Fully-defined saccharide	1437.51	Amphibia Anura Batrachia Bilateria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Lithobates pipiens Lithobates Metazoa Neobatrachia Opisthokonta Rana Ranidae Ranoidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms
G42548XF	GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	G69931GF	Fully-defined saccharide	1275.46	Amphibia Anura Batrachia Bilateria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Lithobates pipiens Lithobates Metazoa Neobatrachia Opisthokonta Rana Ranidae Ranoidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms
G30673AC	GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	G69931GF	Linkage-defined saccharide	1275.46	Amphibia Anura Batrachia Bilateria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Lithobates pipiens Lithobates Metazoa Neobatrachia Opisthokonta Ranidae Ranoidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms
G51688VK	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	G17226QK	Linkage-defined saccharide	1437.51	Amphibia Anura Batrachia Bilateria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Lithobates pipiens Lithobates Metazoa Neobatrachia Opisthokonta Ranidae Ranoidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms

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