GlyCosmos Portal

GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	December 15, 2025

Displaying **all 19** entries
GlyTouCan ID	IUPAC Condensed	Names	Motifs	Archetype	Subsumption Level	Monoisotopic Mass	Organisms
G73001SO	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(a1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	2162.73	Acanthomorphata Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Pollachius pollachius Pollachius Teleostei Teleostomi Vertebrata Zeiogadaria cellular organisms
G40748TG					Linkage-defined saccharide	254.1	Acanthomorphata Actinopteri Actinopterygii Amniota Bacteria Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Ctenosquamata Cyanobacteriota/Melainabacteria group Cyanobacteriota Cyanophyceae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neopterygii Neoteleostei Opisthokonta Oscillatoriaceae Oscillatoriales Oscillatoriophycideae Osteoglossocephalai Paracanthopterygii Phormidium tenue Phormidium Pollachius pollachius Pollachius Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Terrabacteria group Tetrapoda Theria Vertebrata Zeiogadaria cellular organisms
G91237TK	Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	GM3 N-Acetyl GM3	GM3	G40764VP	Fully-defined saccharide	633.21	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Channichthyidae Chinchilla Chinchillidae Chionodraco myersi Chionodraco Chondrichthyes Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Enterobacterales Enterobacteriaceae Epsilonproteobacteria Escherichia coli Escherichia Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Leucoraja erinacea Leucoraja Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Narcinidae Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Oncorhynchus mykiss Oncorhynchus Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Ovis aries Ovis Papio hamadryas Papio Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Pipidae Pipoidea Platyrrhini Primates Protacanthopterygii Pseudocrenilabrinae Pseudomonadota Rajidae Rajiformes Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Trematomus lepidorhinus Trematomus Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus Zeiogadaria cellular organisms
G00031MO	Gal(b1-3)GalNAc(a1-	T Antigen core 1	O-Glycan core 1 T antigen TF antigen	G00032MO	Fully-defined saccharide	383.14	Acalyptratae Acanthomorphata Actinopteri Actinopterygii Amniota Arthropoda Artiodactyla Bacteria Bacteroidaceae Bacteroidales Bacteroides ovatus Bacteroides Bacteroidia Bacteroidota/Chlorobiota group Bacteroidota Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria FCB group Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Neopterygii Neoteleostei Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Panarthropoda Pancrustacea Paracanthopterygii Pecora Pisoniviricetes Pisuviricota Pollachius pollachius Pollachius Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Shigella dysenteriae Shigella Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses Zeiogadaria cellular organisms melanogaster group melanogaster subgroup
G68110IF	Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1a	GT1a	G97898ZO	Fully-defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Bilateria Boreoeutheria Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Catarrhini Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Epsilonproteobacteria Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Primates Pseudocrenilabrinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms
G08648UJ	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G37184KW	Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1b	GD1b	G84052PW	Fully-defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Channichthyidae Chionodraco myersi Chionodraco Chondrichthyes Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Crocidurinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Euacanthomorphacea Euarchontoglires Eukaryota Eulipotyphla Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Leucoraja erinacea Leucoraja Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Narcinidae Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Papio hamadryas Papio Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Pipidae Pipoidea Primates Pseudocrenilabrinae Rajidae Rajiformes Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Soricidae Stenella coeruleoalba Stenella Suidae Suina Suncus murinus Suncus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Xenopodinae Xenopus laevis Xenopus Xenopus Zeiogadaria cellular organisms
G30207PZ	Neu5Ac(a2-3)Gal(b1-	GM4 Neu5Ac(a2-3)Gal(b1)	GM4	G58436AI	Fully-defined saccharide	471.16	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cetacea Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Zeiogadaria cellular organisms
G78818SP	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1c	GT1c	G53140PB	Fully-defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Zeiogadaria cellular organisms
G36476BA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT2	GT2	G66281XG	Fully-defined saccharide	1418.48	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Artiodactyla Bilateria Boreoeutheria Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus morhua Gadus Glires Gnathostomata Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muroidea Myomorpha Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phodopus sungorus Phodopus Pollachius pollachius Pollachius Pseudocrenilabrinae Rodentia Sarcopterygii Scardinius erythrophthalmus Scardinius Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms
G71158TZ	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G08624VL	Neu5Ac9Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-		GD2	G09990WW	Fully-defined saccharide	1460.49	Acanthomorphata Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Pollachius pollachius Pollachius Teleostei Teleostomi Vertebrata Zeiogadaria cellular organisms
G21518JT	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Zeiogadaria cellular organisms
G94547EX	Neu5Ac9Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-		9-O-Acetyl GD3 GD3	G02723HE	Fully-defined saccharide	1257.41	Acanthomorphata Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Pollachius pollachius Pollachius Teleostei Teleostomi Vertebrata Zeiogadaria cellular organisms
G84052PW	Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G15021LG	Glc(?1-			G80753ZP			50 kb inversion clade Acanthomorphata Actinopteri Actinopterygii Amniota Archaea Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euryarchaeota Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Fabaceae Fabales Gadariae Gadidae Gadiformes Gadoidei Glires Gnathostomata Gunneridae Halobacteria Halobacteriales Haloferacaceae Haloferax volcanii Haloferax Haplorrhini Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo IRL clade Magnoliopsida Mammalia Medicago sativa Medicago Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Papilionoideae Paracanthopterygii Pentapetalae Pollachius pollachius Pollachius Primates Rodentia Sarcopterygii Simiiformes Spermatophyta Stenosarchaea group Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Theria Tracheophyta Trifolieae Vertebrata Viridiplantae Zeiogadaria cellular organisms eudicotyledons fabids rosids
G53140PB	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G54434YJ	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc		GM4		Linkage-defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G68158BT	Gal(?1-			G43523TH			Acanthomorphata Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Pollachius pollachius Pollachius Primates Rodentia Sarcopterygii Simiiformes Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Theria Vertebrata Viridiplantae Zeiogadaria cellular organisms

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Last updated: December 8, 2025