GlyCosmos Portal

GlyCosmos Glycoproteins

Integrated list of glycoproteins extracted from UniProt and annotated with glycosylation data from GlyGen and GlyConnect. For each entry, information such as glycosylation site, glycans, diseases, 3D structures, and pathway information are available.

Database	Last Updated
GlyConnect	April 19, 2023
GlycoProtDB	October 1, 2021
GlyGen	October 2, 2023
Human Protein Atlas	June 19, 2023 (Protein Atlas version 23)
MCAW-DB	July 10, 2019
The O-GlcNAc Database	April 10, 2023 (v1.3)
Reactome	February 28, 2024 (Version 87)
UniProt	December 27, 2023

Protein Name	UniProt ID	Gene Symbol	Species	Taxonomy ID	No. of Glycosylation Sites ▼	GO Labels	GO IDs
Protein phosphatase 1 regulatory subunit 12A	O14974	PPP1R12A	Homo sapiens	9606	58	14-3-3 protein binding A band PTW/PP1 phosphatase complex Z disc actin cytoskeleton cellular response to xenobiotic stimulus centrosome centrosome cycle contractile fiber cytoplasm cytosol enzyme inhibitor activity focal adhesion kinetochore mitotic cell cycle negative regulation of catalytic activity nucleolus nucleoplasm phosphatase regulator activity plasma membrane positive regulation of myosin-light-chain-phosphatase activity positive regulation of transcription by RNA polymerase II protein dephosphorylation protein kinase binding regulation of cell adhesion regulation of myosin-light-chain-phosphatase activity regulation of nucleocytoplasmic transport signal transduction	GO:0000278 GO:0000776 GO:0004857 GO:0005654 GO:0005730 GO:0005737 GO:0005813 GO:0005829 GO:0005886 GO:0005925 GO:0006470 GO:0007098 GO:0007165 GO:0015629 GO:0019208 GO:0019901 GO:0030018 GO:0030155 GO:0031672 GO:0035507 GO:0035508 GO:0043086 GO:0043292 GO:0045944 GO:0046822 GO:0071466 GO:0071889 GO:0072357
Agrin	O00468	AGRN	Homo sapiens	9606	58	G protein-coupled acetylcholine receptor signaling pathway Golgi lumen basement membrane calcium ion binding chondroitin sulfate binding clustering of voltage-gated sodium channels collagen-containing extracellular matrix dystroglycan binding extracellular exosome extracellular region heparan sulfate proteoglycan binding laminin binding lysosomal lumen neuromuscular junction development plasma membrane positive regulation of GTPase activity positive regulation of filopodium assembly positive regulation of synaptic assembly at neuromuscular junction positive regulation of transcription by RNA polymerase II receptor clustering sialic acid binding signal transduction structural constituent of cytoskeleton synapse synapse organization	GO:0002162 GO:0005200 GO:0005509 GO:0005576 GO:0005604 GO:0005796 GO:0005886 GO:0007165 GO:0007213 GO:0007528 GO:0033691 GO:0035374 GO:0043113 GO:0043202 GO:0043236 GO:0043395 GO:0043547 GO:0045162 GO:0045202 GO:0045887 GO:0045944 GO:0050808 GO:0051491 GO:0062023 GO:0070062
Neurogenic locus notch homolog protein 1	P46531	NOTCH1	Homo sapiens	9606	57	Golgi membrane MAML1-RBP-Jkappa- ICN1 complex Notch binding Notch signaling involved in heart development Notch signaling pathway Notch signaling pathway involved in regulation of secondary heart field cardioblast proliferation T-helper 17 type immune response acrosomal vesicle adherens junction animal organ regeneration aortic valve morphogenesis apical plasma membrane apoptotic process involved in embryonic digit morphogenesis arterial endothelial cell differentiation astrocyte differentiation atrioventricular node development atrioventricular valve morphogenesis auditory receptor cell fate commitment axon guidance branching morphogenesis of an epithelial tube calcium ion binding calcium-ion regulated exocytosis cardiac atrium morphogenesis cardiac chamber formation cardiac epithelial to mesenchymal transition cardiac left ventricle morphogenesis cardiac muscle cell myoblast differentiation cardiac muscle cell proliferation cardiac muscle tissue morphogenesis cardiac right atrium morphogenesis cardiac right ventricle formation cardiac septum morphogenesis cardiac vascular smooth muscle cell development cardiac ventricle morphogenesis cell differentiation in spinal cord cell migration involved in endocardial cushion formation cell surface cellular response to follicle-stimulating hormone stimulus cellular response to hypoxia cellular response to tumor cell cellular response to vascular endothelial growth factor stimulus chromatin DNA binding cilium assembly collecting duct development compartment pattern specification coronary artery morphogenesis coronary sinus valve morphogenesis coronary vein morphogenesis cytosol determination of left/right symmetry distal tubule development embryonic hindlimb morphogenesis endocardial cell differentiation endocardial cushion morphogenesis endocardium development endocardium morphogenesis endoderm development endoplasmic reticulum membrane enzyme binding enzyme inhibitor activity epidermal cell fate specification epithelial cell fate commitment epithelial cell proliferation epithelial to mesenchymal transition epithelial to mesenchymal transition involved in endocardial cushion formation extracellular region forebrain development foregut morphogenesis glomerular mesangial cell development growth involved in heart morphogenesis hair follicle morphogenesis heart development heart looping heart trabecula morphogenesis homeostasis of number of cells within a tissue humoral immune response identical protein binding immune response in utero embryonic development inflammatory response to antigenic stimulus inhibition of neuroepithelial cell differentiation interleukin-17-mediated signaling pathway keratinocyte differentiation left/right axis specification liver development lung development luteolysis mesenchymal cell development mitral valve formation negative regulation of BMP signaling pathway negative regulation of DNA-templated transcription negative regulation of anoikis negative regulation of biomineral tissue development negative regulation of calcium ion-dependent exocytosis negative regulation of canonical Wnt signaling pathway negative regulation of cardiac muscle hypertrophy negative regulation of catalytic activity negative regulation of cell adhesion molecule production negative regulation of cell migration involved in sprouting angiogenesis negative regulation of cell population proliferation negative regulation of cell proliferation involved in heart valve morphogenesis negative regulation of cell-cell adhesion mediated by cadherin negative regulation of cell-substrate adhesion negative regulation of cold-induced thermogenesis negative regulation of collagen biosynthetic process negative regulation of endothelial cell chemotaxis negative regulation of epithelial cell proliferation negative regulation of extracellular matrix constituent secretion negative regulation of gene expression negative regulation of glial cell proliferation negative regulation of inner ear auditory receptor cell differentiation negative regulation of myoblast differentiation negative regulation of myotube differentiation negative regulation of neurogenesis negative regulation of oligodendrocyte differentiation negative regulation of ossification negative regulation of osteoblast differentiation negative regulation of photoreceptor cell differentiation negative regulation of pro-B cell differentiation negative regulation of stem cell differentiation negative regulation of transcription by RNA polymerase II neural tube development neuroendocrine cell differentiation neuronal stem cell population maintenance nucleoplasm nucleus oligodendrocyte differentiation outflow tract morphogenesis pericardium morphogenesis plasma membrane positive regulation of BMP signaling pathway positive regulation of DNA-templated transcription positive regulation of ERK1 and ERK2 cascade positive regulation of Notch signaling pathway positive regulation of Ras protein signal transduction positive regulation of aorta morphogenesis positive regulation of apoptotic process involved in morphogenesis positive regulation of astrocyte differentiation positive regulation of cardiac epithelial to mesenchymal transition positive regulation of cardiac muscle cell proliferation positive regulation of cell migration positive regulation of cell population proliferation positive regulation of endothelial cell differentiation positive regulation of epithelial cell proliferation positive regulation of gene expression positive regulation of keratinocyte differentiation positive regulation of neuroblast proliferation positive regulation of receptor signaling pathway via JAK-STAT positive regulation of transcription by RNA polymerase II positive regulation of transcription of Notch receptor target positive regulation of viral genome replication prostate gland epithelium morphogenesis protein catabolic process protein import into nucleus pulmonary valve morphogenesis receptor complex regulation of DNA-templated transcription regulation of epithelial cell proliferation involved in prostate gland development regulation of extracellular matrix assembly regulation of somitogenesis regulation of stem cell proliferation regulation of transcription by RNA polymerase II response to lipopolysaccharide response to muramyl dipeptide retinal cone cell differentiation secretory columnal luminar epithelial cell differentiation involved in prostate glandular acinus development skeletal muscle cell differentiation somatic stem cell division spermatogenesis sprouting angiogenesis tissue regeneration transcription by RNA polymerase II transcription coactivator activity transcription regulator activator activity transmembrane signaling receptor activity tube formation vasculogenesis involved in coronary vascular morphogenesis venous endothelial cell differentiation ventricular septum morphogenesis ventricular trabecula myocardium morphogenesis	GO:0000122 GO:0000139 GO:0001554 GO:0001669 GO:0001701 GO:0001837 GO:0001889 GO:0001947 GO:0002040 GO:0002052 GO:0002085 GO:0002193 GO:0002437 GO:0003151 GO:0003157 GO:0003160 GO:0003162 GO:0003169 GO:0003180 GO:0003181 GO:0003182 GO:0003184 GO:0003192 GO:0003198 GO:0003203 GO:0003207 GO:0003208 GO:0003209 GO:0003213 GO:0003214 GO:0003219 GO:0003222 GO:0003241 GO:0003252 GO:0003270 GO:0003273 GO:0003332 GO:0003344 GO:0003713 GO:0004857 GO:0004888 GO:0005112 GO:0005509 GO:0005576 GO:0005634 GO:0005654 GO:0005789 GO:0005829 GO:0005886 GO:0005912 GO:0006355 GO:0006357 GO:0006366 GO:0006606 GO:0006955 GO:0006959 GO:0007219 GO:0007221 GO:0007283 GO:0007368 GO:0007386 GO:0007411 GO:0007440 GO:0007492 GO:0007507 GO:0008284 GO:0008285 GO:0009912 GO:0009957 GO:0009986 GO:0010614 GO:0010628 GO:0010629 GO:0010812 GO:0010832 GO:0014031 GO:0014807 GO:0016324 GO:0017156 GO:0019899 GO:0021515 GO:0021915 GO:0030163 GO:0030216 GO:0030279 GO:0030324 GO:0030335 GO:0030513 GO:0030514 GO:0030900 GO:0031069 GO:0031100 GO:0031490 GO:0032495 GO:0032496 GO:0032966 GO:0035116 GO:0035148 GO:0035914 GO:0035924 GO:0042246 GO:0042670 GO:0042802 GO:0043086 GO:0043235 GO:0045070 GO:0045603 GO:0045608 GO:0045618 GO:0045662 GO:0045668 GO:0045747 GO:0045892 GO:0045893 GO:0045944 GO:0045955 GO:0046427 GO:0046533 GO:0046579 GO:0048103 GO:0048708 GO:0048709 GO:0048711 GO:0048715 GO:0048754 GO:0048873 GO:0050673 GO:0050679 GO:0050680 GO:0050768 GO:0055008 GO:0060038 GO:0060045 GO:0060253 GO:0060271 GO:0060317 GO:0060354 GO:0060379 GO:0060411 GO:0060412 GO:0060528 GO:0060740 GO:0060768 GO:0060842 GO:0060843 GO:0060948 GO:0060956 GO:0060979 GO:0060982 GO:0061101 GO:0061314 GO:0061384 GO:0062043 GO:0070168 GO:0070374 GO:0070986 GO:0071228 GO:0071372 GO:0071456 GO:0072017 GO:0072044 GO:0072091 GO:0072144 GO:0072148 GO:0072538 GO:0090051 GO:0090090 GO:0097150 GO:0097400 GO:0120163 GO:0140537 GO:1901201 GO:1902263 GO:1902339 GO:1903849 GO:2000048 GO:2000737 GO:2000811 GO:2000974 GO:2001027
E1A-binding protein p400	Q96L91	EP400	Homo sapiens	9606	56	ATP binding ATP-dependent chromatin remodeler activity DNA binding DNA repair NuA4 histone acetyltransferase complex Swr1 complex chromatin binding helicase activity hydrolase activity nuclear speck nucleoplasm nucleosome positive regulation of DNA-templated transcription positive regulation of double-strand break repair via homologous recombination protein antigen binding regulation of apoptotic process regulation of cell cycle regulation of double-strand break repair	GO:0000786 GO:0000812 GO:0003677 GO:0003682 GO:0004386 GO:0005524 GO:0005654 GO:0006281 GO:0016607 GO:0016787 GO:0035267 GO:0042981 GO:0045893 GO:0051726 GO:0140658 GO:1905168 GO:1990405 GO:2000779
Nucleoporin p58/p45	Q9BVL2	NUP58	Homo sapiens	9606	56	identical protein binding mRNA transport nuclear envelope nuclear localization sequence binding nuclear membrane nuclear pore nucleocytoplasmic transport protein transport protein-containing complex binding regulation of protein import into nucleus structural constituent of nuclear pore	GO:0005635 GO:0005643 GO:0006913 GO:0008139 GO:0015031 GO:0017056 GO:0031965 GO:0042306 GO:0042802 GO:0044877 GO:0051028
Neurogenic locus notch homolog protein 1	Q01705	Notch1	Mus musculus	10090	55	Golgi membrane MAML1-RBP-Jkappa- ICN1 complex Notch binding Notch signaling involved in heart development Notch signaling pathway Notch signaling pathway involved in regulation of secondary heart field cardioblast proliferation T-helper 17 type immune response acrosomal vesicle adherens junction animal organ regeneration aortic valve morphogenesis apical plasma membrane apoptotic process apoptotic process involved in embryonic digit morphogenesis arterial endothelial cell differentiation astrocyte differentiation atrioventricular node development atrioventricular valve morphogenesis auditory receptor cell fate commitment axon guidance axonogenesis branching morphogenesis of an epithelial tube calcium ion binding calcium-ion regulated exocytosis cardiac atrium morphogenesis cardiac chamber formation cardiac epithelial to mesenchymal transition cardiac left ventricle morphogenesis cardiac muscle cell myoblast differentiation cardiac muscle cell proliferation cardiac muscle tissue morphogenesis cardiac right atrium morphogenesis cardiac right ventricle formation cardiac septum morphogenesis cardiac vascular smooth muscle cell development cardiac ventricle morphogenesis cell differentiation cell differentiation in spinal cord cell migration involved in endocardial cushion formation cell periphery cell population proliferation cell surface cellular response to follicle-stimulating hormone stimulus cellular response to hypoxia cellular response to tumor cell cellular response to vascular endothelial growth factor stimulus chromatin DNA binding chromatin binding cilium assembly collecting duct development compartment pattern specification coronary artery morphogenesis coronary sinus valve morphogenesis coronary vein morphogenesis cytoplasm cytoplasmic vesicle cytoskeleton cytosol determination of left/right symmetry distal tubule development embryonic hindlimb morphogenesis embryonic limb morphogenesis endocardial cell differentiation endocardial cushion development endocardial cushion morphogenesis endocardium development endocardium morphogenesis endoderm development endoplasmic reticulum endoplasmic reticulum membrane endosome membrane enzyme binding enzyme inhibitor activity epidermal cell fate specification epidermis development epithelial cell fate commitment epithelial cell proliferation epithelial to mesenchymal transition epithelial to mesenchymal transition involved in endocardial cushion formation extracellular region forebrain development foregut morphogenesis gene expression glial cell differentiation glomerular mesangial cell development growth involved in heart morphogenesis hair follicle morphogenesis heart development heart looping heart trabecula morphogenesis homeostasis of number of cells within a tissue humoral immune response identical protein binding in utero embryonic development inflammatory response to antigenic stimulus inhibition of neuroepithelial cell differentiation interleukin-17-mediated signaling pathway keratinocyte differentiation lamellipodium left/right axis specification liver development lung development luteolysis mesenchymal cell development mitral valve formation negative regulation of BMP signaling pathway negative regulation of DNA-templated transcription negative regulation of anoikis negative regulation of biomineral tissue development negative regulation of calcium ion-dependent exocytosis negative regulation of canonical Wnt signaling pathway negative regulation of cardiac muscle hypertrophy negative regulation of catalytic activity negative regulation of cell adhesion molecule production negative regulation of cell migration involved in sprouting angiogenesis negative regulation of cell population proliferation negative regulation of cell proliferation involved in heart valve morphogenesis negative regulation of cell-cell adhesion mediated by cadherin negative regulation of cell-substrate adhesion negative regulation of cold-induced thermogenesis negative regulation of collagen biosynthetic process negative regulation of endothelial cell chemotaxis negative regulation of epithelial cell proliferation negative regulation of extracellular matrix constituent secretion negative regulation of gene expression negative regulation of glial cell proliferation negative regulation of inner ear auditory receptor cell differentiation negative regulation of myoblast differentiation negative regulation of myotube differentiation negative regulation of neurogenesis negative regulation of neuron differentiation negative regulation of oligodendrocyte differentiation negative regulation of ossification negative regulation of osteoblast differentiation negative regulation of photoreceptor cell differentiation negative regulation of pro-B cell differentiation negative regulation of programmed cell death negative regulation of stem cell differentiation negative regulation of transcription by RNA polymerase II neural tube development neuroendocrine cell differentiation neuron differentiation neuron fate commitment neuronal stem cell population maintenance nucleoplasm nucleus oligodendrocyte differentiation osteoblast fate commitment outflow tract morphogenesis pericardium morphogenesis plasma membrane positive regulation of BMP signaling pathway positive regulation of DNA-templated transcription positive regulation of ERK1 and ERK2 cascade positive regulation of Notch signaling pathway positive regulation of Ras protein signal transduction positive regulation of aorta morphogenesis positive regulation of apoptotic process positive regulation of apoptotic process involved in morphogenesis positive regulation of astrocyte differentiation positive regulation of cardiac epithelial to mesenchymal transition positive regulation of cardiac muscle cell proliferation positive regulation of cell migration positive regulation of cell population proliferation positive regulation of endothelial cell differentiation positive regulation of ephrin receptor signaling pathway positive regulation of epithelial cell proliferation positive regulation of epithelial to mesenchymal transition positive regulation of gene expression positive regulation of glial cell differentiation positive regulation of keratinocyte differentiation positive regulation of neuroblast proliferation positive regulation of receptor signaling pathway via JAK-STAT positive regulation of transcription by RNA polymerase II positive regulation of transcription of Notch receptor target positive regulation of viral genome replication prostate gland epithelium morphogenesis protein catabolic process protein import into nucleus pulmonary valve morphogenesis receptor complex regulation of Notch signaling pathway regulation of cardioblast proliferation regulation of cell adhesion involved in heart morphogenesis regulation of cell migration regulation of cell population proliferation regulation of epithelial cell proliferation regulation of epithelial cell proliferation involved in prostate gland development regulation of extracellular matrix assembly regulation of gene expression regulation of inner ear auditory receptor cell differentiation regulation of neurogenesis regulation of somitogenesis regulation of stem cell proliferation regulation of transcription by RNA polymerase II response to lipopolysaccharide response to muramyl dipeptide retinal cone cell differentiation ruffle secretory columnal luminar epithelial cell differentiation involved in prostate glandular acinus development skeletal muscle cell differentiation somatic stem cell division spermatogenesis sprouting angiogenesis tissue regeneration transcription by RNA polymerase II transcription coactivator activity transcription regulator activator activity transmembrane signaling receptor activity tube formation vasculogenesis involved in coronary vascular morphogenesis venous blood vessel morphogenesis venous endothelial cell differentiation ventricular septum morphogenesis ventricular trabecula myocardium morphogenesis	GO:0000122 GO:0000139 GO:0001554 GO:0001669 GO:0001701 GO:0001726 GO:0001837 GO:0001889 GO:0001947 GO:0002040 GO:0002051 GO:0002052 GO:0002085 GO:0002193 GO:0002437 GO:0003151 GO:0003157 GO:0003160 GO:0003162 GO:0003169 GO:0003180 GO:0003181 GO:0003182 GO:0003184 GO:0003192 GO:0003197 GO:0003198 GO:0003203 GO:0003207 GO:0003208 GO:0003209 GO:0003213 GO:0003214 GO:0003219 GO:0003222 GO:0003241 GO:0003252 GO:0003264 GO:0003270 GO:0003273 GO:0003332 GO:0003344 GO:0003682 GO:0003713 GO:0004857 GO:0004888 GO:0005112 GO:0005509 GO:0005576 GO:0005634 GO:0005654 GO:0005737 GO:0005783 GO:0005789 GO:0005829 GO:0005856 GO:0005886 GO:0005912 GO:0006357 GO:0006366 GO:0006606 GO:0006915 GO:0006959 GO:0007219 GO:0007221 GO:0007283 GO:0007368 GO:0007386 GO:0007409 GO:0007411 GO:0007440 GO:0007492 GO:0007507 GO:0008283 GO:0008284 GO:0008285 GO:0008544 GO:0008593 GO:0009912 GO:0009957 GO:0009986 GO:0010001 GO:0010008 GO:0010467 GO:0010468 GO:0010614 GO:0010628 GO:0010629 GO:0010718 GO:0010812 GO:0010832 GO:0014031 GO:0014807 GO:0016324 GO:0017156 GO:0019899 GO:0021515 GO:0021915 GO:0030027 GO:0030154 GO:0030163 GO:0030182 GO:0030216 GO:0030279 GO:0030324 GO:0030326 GO:0030334 GO:0030335 GO:0030513 GO:0030514 GO:0030900 GO:0031069 GO:0031100 GO:0031410 GO:0031490 GO:0032495 GO:0032496 GO:0032966 GO:0035116 GO:0035148 GO:0035914 GO:0035924 GO:0042127 GO:0042246 GO:0042670 GO:0042802 GO:0043065 GO:0043069 GO:0043086 GO:0043235 GO:0045070 GO:0045603 GO:0045607 GO:0045608 GO:0045618 GO:0045662 GO:0045665 GO:0045668 GO:0045687 GO:0045747 GO:0045892 GO:0045893 GO:0045944 GO:0045955 GO:0046427 GO:0046533 GO:0046579 GO:0048103 GO:0048663 GO:0048708 GO:0048709 GO:0048711 GO:0048715 GO:0048754 GO:0048845 GO:0048873 GO:0050673 GO:0050678 GO:0050679 GO:0050680 GO:0050767 GO:0050768 GO:0055008 GO:0060038 GO:0060045 GO:0060253 GO:0060271 GO:0060317 GO:0060354 GO:0060379 GO:0060411 GO:0060412 GO:0060528 GO:0060740 GO:0060768 GO:0060842 GO:0060843 GO:0060948 GO:0060956 GO:0060979 GO:0060982 GO:0061101 GO:0061314 GO:0061344 GO:0061384 GO:0062043 GO:0070168 GO:0070374 GO:0070986 GO:0071228 GO:0071372 GO:0071456 GO:0071944 GO:0072017 GO:0072044 GO:0072091 GO:0072144 GO:0072148 GO:0072538 GO:0090051 GO:0090090 GO:0097150 GO:0097400 GO:0120163 GO:0140537 GO:1901189 GO:1901201 GO:1902263 GO:1902339 GO:1903849 GO:2000048 GO:2000737 GO:2000811 GO:2000974 GO:2001027
Spike glycoprotein	P0DTC2	S	Severe acute respiratory syndrome coronavirus 2	2697049	54	fusion of virus membrane with host endosome membrane fusion of virus membrane with host plasma membrane host cell endoplasmic reticulum-Golgi intermediate compartment membrane host cell plasma membrane host cell surface host cell surface receptor binding host extracellular space identical protein binding membrane fusion plasma membrane positive regulation of viral entry into host cell receptor-mediated endocytosis of virus by host cell receptor-mediated virion attachment to host cell structural constituent of virion suppression by virus of host tetherin activity suppression by virus of host type I interferon-mediated signaling pathway viral envelope viral translation virion attachment to host cell virion membrane	GO:0005886 GO:0019031 GO:0019062 GO:0019064 GO:0019065 GO:0019081 GO:0020002 GO:0039502 GO:0039587 GO:0039654 GO:0039660 GO:0042802 GO:0043655 GO:0044173 GO:0044228 GO:0046598 GO:0046789 GO:0046813 GO:0055036 GO:0061025
Foot protein 1 variant 2	A1X159	fp-1	Perna viridis	73031	53	extracellular region	GO:0005576
Cysteine-rich, acidic integral membrane protein	Q03650	CRAM	Trypanosoma brucei brucei	5702	53	ciliary pocket endocytosis plasma membrane	GO:0005886 GO:0006897 GO:0020016
Bud site selection protein RAX2	Q12465	RAX2	Saccharomyces cerevisiae S288C	559292	52	axial cellular bud site selection bipolar cellular bud site selection cellular bud neck cellular bud scar cellular bud site selection cellular bud tip endoplasmic reticulum mitochondrion plasma membrane of growing cell tip protein localization	GO:0000282 GO:0005621 GO:0005739 GO:0005783 GO:0005934 GO:0005935 GO:0007120 GO:0007121 GO:0008104 GO:1902929