GlyCosmos Portal

Annotation

Keyword

Disulfide bond
Glycoprotein
Glycosyltransferase
Golgi apparatus
Lipid metabolism
Manganese
Metal-binding
Reference proteome
Secreted
Signal-anchor
Transmembrane helix

Proteins

Displaying 1 entry
UniProt	Protein Name
Q9JJ04	Beta-N-acetylglucosaminyl-glycolipid beta-1,4-galactosyltransferase Lactotriaosylceramide beta-1,4-galactosyltransferase N-acetyllactosamine synthase Nal synthase UDP-Gal:beta-GlcNAc beta-1,4-galactosyltransferase 4 UDP-galactose:beta-N-acetylglucosamine beta-1,4-galactosyltransferase 4

Annotation information from UniProt.

Gene Ontology (GO)

Displaying **all 5** entries
GO Term	Evidence Code	PMID
protein glycosylation	IEA
lactosylceramide biosynthetic process	ISO ISS
keratan sulfate biosynthetic process	ISO ISS
carbohydrate metabolic process	IEA
glycosylation	IBA

GO Hierarchy

biological process

metabolic process [inference]

nitrogen compound metabolic process [inference]

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

amide metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

biosynthetic process [inference]

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organic substance biosynthetic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular metabolic process [inference]

sulfur compound metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

primary metabolic process [inference]

carbohydrate metabolic process

lipid metabolic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

glycosylation

macromolecule glycosylation [inference]

protein glycosylation

organic substance metabolic process [inference]

carbohydrate metabolic process

lipid metabolic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

macromolecule modification [inference]

protein modification process [inference]

protein glycosylation

macromolecule glycosylation [inference]

protein glycosylation

carbohydrate derivative metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organic substance biosynthetic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular process [inference]

cellular metabolic process [inference]

sulfur compound metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

Displaying **all 3** entries
GO Term	Evidence Code	PMID
Golgi apparatus	IBA ISO
extracellular region	IEA
Golgi membrane	ISO ISS

GO Hierarchy

cellular component

cellular anatomical entity [inference]

extracellular region

membrane [inference]

organelle membrane [inference]

bounding membrane of organelle [inference]

Golgi membrane

organelle [inference]

membrane-bounded organelle [inference]

intracellular membrane-bounded organelle [inference]

Golgi apparatus

intracellular organelle [inference]

intracellular membrane-bounded organelle [inference]

Golgi apparatus

Displaying **all 4** entries
GO Term	Evidence Code	PMID
UDP-galactosyltransferase activity	ISO ISS
metal ion binding	IEA
N-acetyllactosamine synthase activity	ISO ISS
galactosyltransferase activity	IBA

GO Hierarchy

molecular function

catalytic activity [inference]

transferase activity [inference]

glycosyltransferase activity [inference]

UDP-glycosyltransferase activity [inference]

UDP-galactosyltransferase activity

N-acetyllactosamine synthase activity

hexosyltransferase activity [inference]

galactosyltransferase activity

UDP-galactosyltransferase activity

N-acetyllactosamine synthase activity

binding [inference]

ion binding [inference]

cation binding [inference]

metal ion binding

Gene Ontology information from NCBI Gene and UniProt.

OrthoDB (Group)

Group level

Rodentia

Group Name

Beta-1,4-galactosyltransferase

Functional Category

G: Carbohydrate transport and metabolism
I: Lipid transport and metabolism
M: Cell wall/membrane/envelope biogenesis
O: Posttranslational modification, protein turnover, chaperones

Displaying **all 4** entries
Gene Ontology
galactosyltransferase activity
glycosylation
metal ion binding
transferase activity

Displaying **all 4** entries
InterPro
Beta-1,4-galactosyltransferase
Galactosyltransferase, C-terminal
Galactosyltransferase, N-terminal
Nucleotide-diphospho-sugar transferases

KEGG BRITE Database

Orthology

K07969

Name

beta-1,4-galactosyltransferase 4 [EC:2.4.1.90 2.4.1.275 2.4.1.-]

References

9792633

Reactions

Displaying **all 3** entries
KEGG Reaction	Enzyme	Acceptor	Product
R07617	2.4.1.-	G00033MO	G64973KT
R05977	2.4.1.- 2.4.1.275	G06239NA	G58896AZ
R06033	2.4.1.-	G70874WW	G84034JH

Rhea

RHEA:31499

a β-D-GlcNAc-(1→3)-β-D-Gal-(1→4)-β-D-Glc-(1↔1)-Cer(d18:1(4E)) + UDP-α-D-galactose

H⁺ + a neolactoside nLc4Cer(d18:1(4E)) + UDP

Enzyme

2.4.1.275

PMID

RHEA:17745

N-acetyl-D-glucosamine + UDP-α-D-galactose

H⁺ + UDP + β-D-galactosyl-(1→4)-N-acetyl-D-glucosamine

Enzyme

2.4.1.90

PMID

LIPID MAPS Gene / Proteome Database

LMPD ID: LMP004132
Gene Name: UDP-Gal:betaGlcNAc beta 1,4-galactosyltransferase, polypeptide 4

Ortholog

Displaying entries **121 - 124** of **124** in total

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Species	Gene ID	OrthoDB
Peromyscus californicus insignis	128112030	564181_0:00135b
Psammomys obesus	129689628	48139_1:00390d
Chionomys nivalis	130872247	269649_0:000e29
Peromyscus eremicus	131922337	42410_0:00397c

GlyTouCan

GlyComb

GlycoPOST

UniCarb-DR

LM-GlycoRepo

All Resources

Other Information

Tools

Search Tools

Summary

External Links

Annotation

OrthoDB (Group)

KEGG BRITE Database

LIPID MAPS Gene / Proteome Database

International Collaboration

Acknowledgements