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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	February 17, 2025

Displaying entries **251 - 275** of **1270** in total

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GlyTouCan ID ▲	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G22625SJ			Base composition	2281.85	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G22658BN	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)[Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)]Man(a1-3)[Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage-defined saccharide	2734.99	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G22725MK	Gal(?1-?)[Fuc(?1-?)]GlcNAc(?1-		Topology	529.2	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G22754FQ	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	2370.86	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G22768VO	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Fully-defined saccharide	910.33	50 kb inversion clade Acalyptratae Amniota Amphibia Anura Arthropoda Artiodactyla Batrachia Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Neobatrachia Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rana Ranidae Ranoidea Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids
G23000DQ	GlcNAc(b1-6)Man(a1-6)Man(b1-	Mannose O-Mannose	Fully-defined saccharide	545.2	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23010ZW			Monosaccharide composition	3026.09	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23119MJ	Fuc(a1-2)Gal(b1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	H antigen (type 3) N-acetyllactosamine O-Glycan core 2 Type 2 LN	Fully-defined saccharide	1040.39	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23182LG	Neu5Gc(?2-?)Gal(?1-?)GlcNAc(?1-?)[Neu5Gc(?2-?)Gal(?1-?)GlcNAc(?1-?)]Man(a1-3)[Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage-defined saccharide	2912.02	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23232XI	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage-defined saccharide	1519.53	Amniota Articulavirales Artverviricota Ascomycota Aspergillaceae Aspergillus oryzae Aspergillus subgen. Circumdati Aspergillus Avulavirinae Bilateria Boreoeutheria Caniformia Carnivora Catarrhini Chordata Craniata Cricetidae Cricetinae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Filoviridae Fowl plague virus Friend mink cell focus-forming virus Friend murine leukemia virus Friend spleen focus-forming virus Fungi Gammaretrovirus Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Insthoviricetes Laurasiatheria Lentivirus Mammalia Marburgvirus Metazoa Monjiviricetes Mononegavirales Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Mustela lutreola Mustela Mustelidae Mustelinae Musteloidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthoavulavirus javaense Orthoavulavirus Orthomyxoviridae Orthoretrovirinae Orthornavirae Paramyxoviridae Pararnavirae Pezizomycotina Polyploviricotina Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses avian paramyxovirus 1 cellular organisms leotiomyceta saccharomyceta unclassified Gammaretrovirus unclassified Orthomyxoviridae unidentified influenza virus
G23294PN			Monosaccharide composition	1890.66	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G23347CS	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-3)Gal(b1-4)Glc(b1-	GM4 Lactotriaosylceramide Lc3 Type 1 LN	Fully-defined saccharide	1289.44	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23368XB	Gal(?1-?)[Fuc(?1-?)]GlcNAc(?1-?)[Gal(?1-?)]GalNAc(?1-		Topology	894.33	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23399NX	Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)Man(a1-3)[GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage-defined saccharide	1770.66	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23432EQ			Base composition	1681.62	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G23453IV			Base composition	1275.46	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G23616ZX	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Fully-defined saccharide	2879.01	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ovis aries Ovis Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23627AX	Neu5Gc(?2-?)Neu5Ac(?2-?)GalNAc(?1-		Topology	819.28	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23707SD	Gal(b1-4)GlcNAc(b1-2)Man	N-acetyllactosamine Type 2 LN	Linkage-defined saccharide	545.2	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23817OE	GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage-defined saccharide	1640.59	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G23863VK			Base composition	1786.65	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G24000JV	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)[Gal(?1-?)GlcNAc(?1-?)]Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage-defined saccharide	2458.87	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24034CH	Neu5Gc(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	Asialo-GM1 N-Glycolyl-GM2	Fully-defined saccharide	1014.34	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli K-12 Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Pseudomonadota Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24132CP		Lewis x Lex N-Glycan core basic SSEA-1	Linkage-defined saccharide	2443.9	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24393WU	Man(a1-2)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc		Linkage-defined saccharide	910.33	50 kb inversion clade Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Rodentia Sarcopterygii Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Vigna radiata var. radiata Vigna radiata Vigna Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids