GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **526 - 550** of **1011** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▲	Monoisotopic Mass	Organisms
G61764UU			Linkage defined saccharide	1027.29	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G61924UW	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Gc(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2238.78	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G61941WO	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G62167DK	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	912.34	Aculeata Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Anthophila Apidae Apocrita Apoidea Archelosauria Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Primates Protostomia Pterygota Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G62387ZO	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	2222.78	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G62968IF	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2516.91	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthomarburgvirus marburgense Orthomarburgvirus Orthoretrovirinae Orthornavirae Pararnavirae Pecora Primate lentivirus group Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G63035DM	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Amniota Articulavirales Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Discoba Embryophyta Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insthoviricetes Kinetoplastea Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Mesocricetus auratus Mesocricetus Metakinetoplastina Metazoa Muroidea Myomorpha NPAAA clade Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Polyploviricotina Primates Riboviria Rodentia Ruminantia Sarcopterygii Schizotrypanum Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G63065ZE	Neu5Ac(a2-6)Gal(b1-4)[Neu5Ac(a2-6)]GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3170.11	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G62979KP	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	1355.46	Adenoviridae Amniota Artiodactyla Artverviricota Bamfordvirae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend murine leukemia virus Gammaretrovirus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human adenovirus 2 Human mastadenovirus C Laurasiatheria Mammalia Mastadenovirus Metazoa Murine leukemia virus Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Pecora Preplasmiviricota Primates Retroviridae Revtraviricetes Riboviria Rowavirales Ruminantia Sarcopterygii Simiiformes Tectiliviricetes Teleostomi Tetrapoda Theria Varidnaviria Vertebrata Viruses cellular organisms
G63150NE	Gal6S(b1-4)GlcNAc6S(b1-3)Gal6S(b1-4)GlcNAc6S-ol		Linkage defined saccharide	1070.12	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63240YA	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Sialylated LDN Type 2 LN	Linkage defined saccharide	2265.83	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63452MH	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-3)[GlcNAc(b1-6)]GalNAc-ol	Blood group A (type 2)	Linkage defined saccharide	1140.45	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63413QV	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2587.92	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G63617UP	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	2005.72	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63748AU	Neu5Ac(a2-3)Gal(b1-3)[Gal6S(b1-4)GlcNAc6S(b1-3)Gal(b1-4)GlcNAc6S(b1-6)]GalNAc-ol	GM4	Linkage defined saccharide	1646.39	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63821SF	Neu5Gc(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2255.79	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64085NR	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3/6)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2518.93	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64183CB	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc(b1-	GM1b cisGM1	Linkage defined saccharide	1201.42	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Pipidae Pipoidea Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus cellular organisms
G64311TT	Fuc(a1-3)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-3)GlcNAc(b1-2)Man(a1-6)]Man(?1-	Mannose O-Mannose	Linkage defined saccharide	1202.44	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64330NA	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1890.66	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64208CW	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(?1-	Mannose N-acetyllactosamine O-Mannose Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64894MA	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	I' antigen N-Glycan complex Polysialic acid Type 2 LN2	Linkage defined saccharide	4411.56	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Mesocricetus auratus Mesocricetus Metazoa Muroidea Myomorpha Opisthokonta Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64919UI	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM4 LcGg4 Type 1 LN	Linkage defined saccharide	1404.5	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G65414VM	Gal(b1-4)[Fuc(a1-3)]GlcNAc6S-ol		Linkage defined saccharide	611.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G65415JG	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms