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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **5826 - 5850** of **7308** in total

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GlyTouCan ID ▼	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G19829VK	Man(a1-6)Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc		Linkage defined saccharide	894.33	Aculeata Amniota Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dicondylia Dipnotetrapodomorpha Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Hexapoda Hymenoptera Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19822MG	GlcA3S(b1-3)Gal(b1-3)[Neu5Ac(a2-6)]Gal(b1-4)Xyl(?1-		Linkage defined saccharide	1021.24	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19813SH		O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	820.3	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19763QB	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	2046.71	Amniota Ascomycota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Schizosaccharomyces pombe Schizosaccharomyces Schizosaccharomycetaceae Schizosaccharomycetales Schizosaccharomycetes Simiiformes Taphrinomycotina Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19749DC	Gal(b1-3)GlcNAc(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 3 Sialyl Tn antigen Type 1 LN	Linkage defined saccharide	877.32	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19747OA	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1396.49	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acer pseudoplatanus Acer Acereae Aculeata Aesculus hippocastanum Aesculus Amniota Amphiesmenoptera Amphipyrinae Anthemideae Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus awamori Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asterias rubens Asterias Asteriidae Asteroidea Asteroideae Asterozoa Aves BOP clade Batoidea Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Brassicales CUG-Ser1 clade Candida albicans Candida/Lodderomyces clade Candida Caprinae Carica papaya Carica Caricaceae Carpinus betulus Carpinus Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chondrichthyes Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Debaryomycetaceae Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Echinodermata Elasmobranchii Eleutherozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Forcipulatacea Forcipulatida Friend murine leukemia virus Fungi Galliformes Galloanserae Gallus gallus Gallus Gammaretrovirus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera IRL clade Insecta Insthoviricetes Kinetoplastea Komagataella pastoris Komagataella Lamiales Laurasiatheria Lepidoptera Liliopsida Macaca fascicularis Macaca Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Mesocricetus auratus Mesocricetus Metakinetoplastina Metazoa Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Ortervirales Orthomyxoviridae Orthoretrovirinae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Ovis aries Ovis Panarthropoda Pancrustacea Papilionoideae Pararnavirae Pecora Pentapetalae Petrosaviidae Pezizomycotina Phaffomycetaceae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Phasianinae Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Retroviridae Revtraviricetes Riboviria Ricinus communis Ricinus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Santalales Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Schizotrypanum Simiiformes Solanaceae Solanales Sophophora Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma cruzi Trypanosoma dionisii Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses Viscaceae Viscum album Viscum artificial sequences asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup other entries other sequences rosids saccharomyceta synthetic construct unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G19710JA	Gal(b1-3/4)[Fuc(a1-3/4)]GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Lea Lewis a Lewis x Lex N-acetyllactosamine SSEA-1 Type 1 LN Type 2 LN	Linkage defined saccharide	1772.67	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19687OL	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	I' antigen N-Glycan complex	Linkage defined saccharide	3101.12	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19675BE	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)[GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GalNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Blood group A (type 2) Blood group H (type 2) I' antigen Paragloboside	Linkage defined saccharide	2500.92	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19620VP	Gal(a1-3)[Fuc(a1-2)]Gal(b1-3)GalNAc(b1-3)Gal(b1-4)Glc(b1-	Lactosylceramide a-Gal antigen	Linkage defined saccharide	1015.36	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19619KS	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-	SDLex	Linkage defined saccharide	1331.49	Amniota Bilateria Boreoeutheria Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Macaca fuscata Macaca mulatta Macaca Mammalia Metazoa Muroidea Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19615BB	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2240.82	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19607EP	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	3535.24	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19603RR			Glycosidic topology	3085.13	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G19600YQ	Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1745.62	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19594VJ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(?1-4)GlcNAc(b1-4)GlcNAc	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2005.72	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19588HA	Gal(b1-?)GlcNAc(b1-2)[Gal(b1-3)GlcNAc(b1-4)]Man(a1-?)[Gal(b1-?)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	2151.78	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19577LJ	GlcN(b1-				Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19555CW	Neu5Ac(a2-3)Gal(b1-3)[Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	Blood group H (type 1) GM4 I' antigen Leb Lewis b Type 2 LN2	Linkage defined saccharide	2063.77	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19541IU	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	I' antigen N-Glycan complex Polysialic acid Type 2 LN2	Linkage defined saccharide	4046.43	Amniota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muroidea Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19537TS	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19534EE	Gal(b1-3)GlcNAc(b1-4)[Fuc(a1-2)Gal(b1-6)]GalNAc-ol	Type 1 LN	Linkage defined saccharide	896.35	Amniota Bilateria Boreoeutheria Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Macaca radiata Macaca Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19524AS	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	691.25	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19521BV	Man(a1-?)Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)][Fuc(a1-6)]GlcNAc(b1-		Linkage defined saccharide	1040.39	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G19520XO	Gal(a1-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2434.86	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms