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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **6601 - 6625** of **9408** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▲	Monoisotopic Mass	Organisms
G84053BQ	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	1464.56	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend spleen focus-forming virus Gammaretrovirus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Oryctolagus cuniculus Oryctolagus Pararnavirae Pecora Perissodactyla Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Gammaretrovirus
G84170WO	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1722.61	50 kb inversion clade Acalyptratae Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Ascomycota Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Chordata Coelurosauria Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Galliformes Galloanserae Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Influenza A virus Insecta Insthoviricetes Kinetoplastea Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neoptera Opisthokonta Orthomyxoviridae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unidentified influenza virus
G84034JH	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	I' antigen Paragloboside Type 2 LN2	Linkage defined saccharide	1072.38	Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Betaproteobacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Cavia porcellus Cavia Caviidae Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muroidea Myomorpha Neisseria gonorrhoeae Neisseria Neisseriaceae Neisseriales Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Pseudomonadota Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G84064AY	GlcNAc(b1-2)[GlcNAc(b1-2)Man(a1-3)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1640.59	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G84153KU	Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2354.86	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84186YN	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3681.3	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84020NP	Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage defined saccharide	1033.37	Amniota Artiodactyla Avulavirinae Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Orthoavulavirus javaense Orthoavulavirus Orthornavirae Paramyxoviridae Primates Rattus Riboviria Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses avian paramyxovirus 1 cellular organisms
G84167AV	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Sialylated LDN Type 2 LN	Linkage defined saccharide	2263.81	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84147VQ	Gal6S(b1-4)GlcNAc(b1-3)GalNAc(a1-	O-Glycan core 3	Linkage defined saccharide	666.18	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84168AG	GlcA3S(b1-3)Gal(b1-3)[Neu5Ac(a2-6)]Gal(b1-4)Xyl(a1-		Linkage defined saccharide	1021.24	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84155GY		N-Glycan core basic	Linkage defined saccharide	1380.49	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84219QP	Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-		Linkage defined saccharide	869.3	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84224TW	Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	342.12	Amniota Asteroidea Asterozoa Bacteria Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Discoba Echinodermata Eleutherozoa Enterobacterales Enterobacteriaceae Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kinetoplastea Klebsiella sp. Klebsiella/Raoultella group Klebsiella Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Mammalia Metakinetoplastina Metazoa Opisthokonta Primates Pseudomonadota Salmonella Sarcopterygii Shigella flexneri Shigella Simiiformes Teleostomi Tetrapoda Theria Trypanosomatida Trypanosomatidae Vertebrata cellular organisms unclassified Klebsiella
G84260SO	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2046.75	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Glires Gnathostomata Mammalia Meleagridinae Meleagris gallopavo Meleagris Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G84263XI	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-3)Gal(a1-	GM4	Linkage defined saccharide	836.29	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84287MH	Neu5Ac(a2-6)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Sialylated LDN Type 2 LN	Linkage defined saccharide	1972.71	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84316BO	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2032.77	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84301XG	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	I' antigen Type 2 LN2	Linkage defined saccharide	3249.19	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84330VA	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex VIM-2	Linkage defined saccharide	4192.49	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84440YT	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1845.69	Amniota Articulavirales Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Capra hircus Capra Caprinae Carnivora Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insthoviricetes Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Polyploviricotina Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G84448IT	Man(a1-6)Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol		Linkage defined saccharide	896.35	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84350SH	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)GlcNAc(?1-	3-Sialyl-LN (type 2)	Linkage defined saccharide	965.33	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84368MT	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	I' antigen Lewis x Lex N-Glycan complex SSEA-1 Type 2 LN2	Linkage defined saccharide	2151.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84390MS	Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	I' antigen N-Glycan complex Polysialic acid Type 2 LN2	Linkage defined saccharide	4120.47	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G84406HL	Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	LcGg4 Paragloboside a-Gal antigen	Linkage defined saccharide	1421.52	Acanthomorphata Actinopteri Actinopterygii Bilateria Carangaria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Parophrys vetulus Parophrys Percomorphaceae Pleuronectidae Pleuronectiformes Pleuronectoidei Teleostei Teleostomi Vertebrata cellular organisms