GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1176 - 1200** of **9190** in total

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GlyTouCan ID ▼	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G86475BX	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	I' antigen Lewis x Lex N-Glycan complex SSEA-1 Type 2 LN2	Linkage defined saccharide	2442.88	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86469VE	Neu5Ac(a2-3)Gal(b1-3)GalNAc(a1-4)Gal(b1-3)GalNAc(?1-	GM4 O-Glycan core 1 fuzzy	Linkage defined saccharide	1039.37	Amphibia Anura Batrachia Bilateria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Lithobates palustris Lithobates Metazoa Neobatrachia Opisthokonta Ranidae Ranoidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms
G86459PH	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage defined saccharide	1195.42	Amniota Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus oryzae Aspergillus subgen. Circumdati Aspergillus Avulavirinae Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Cricetidae Cricetinae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Friend murine leukemia virus Fungi Gammaretrovirus Glires Gnathostomata Haploviricotina Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthoavulavirus javaense Orthoavulavirus Orthoretrovirinae Orthornavirae Ovis aries Ovis Paramyxoviridae Pararnavirae Pecora Pezizomycotina Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses avian paramyxovirus 1 cellular organisms leotiomyceta saccharomyceta
G86451IJ					Amniota Anacardiaceae Anacardium occidentale Anacardium Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Magnoliopsida Mammalia Mesangiospermae Metazoa Opisthokonta Pentapetalae Primates Sapindales Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids
G86442DH		LDN LacdiNAc	Linkage defined saccharide	1316.49	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86439QE	Neu5Ac(a2-6)Gal(b1-4)GlcNAc6S(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	2,6-sialyl-Sulfo-LN 6-sialyl-6-sulfo LN N-Glycan core basic N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2319.76	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86433UB	Neu5Ac8S(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	916.25	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86419OL	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-		Linkage defined saccharide	1517.51	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86414TE		I' antigen N-Glycan complex Type 2 LN2	Linkage defined saccharide	3392.22	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86414FL	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1934.72	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86408JD			Glycosidic topology	2614.96	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G86399IJ	Man(a1-3)[Man(a1-3/6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1072.38	Amniota Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muroidea Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86388AX	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	I' antigen Paragloboside Polysialic acid Type 2 LN2	Linkage defined saccharide	1654.57	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86387VT	Fuc(a1-2)Gal(b1-?)GlcNAc(b1-3)Gal(b1-3)[Gal(b1-6)]GalNAc(a1-	Blood group H (type 1) Blood group H (type 2) O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	1056.39	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86386QT	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1275.46	Amniota Amphibia Anas platyrhynchos Anas sp. Anas Anatidae Anatinae Anseriformes Anura Archelosauria Archosauria Aves Batrachia Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Leporidae Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Perdicinae Phasianidae Phasianinae Primates Ranidae Ranoidea Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified Anas unclassified entries unclassified sequences unidentified
G86385RC	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Alpharhabdovirinae Amniota Amphiesmenoptera Amphipyrinae Anserineae Anthemideae Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae Aves BOP clade Batoidea Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Carpinus betulus Carpinus Caryophyllales Catarrhini Cavia porcellus Cavia Caviidae Chenopodiaceae Chenopodioideae Chondrichthyes Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Fungi Galliformes Galloanserae Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha IRL clade Insecta Insthoviricetes Kinetoplastea Komagataella pastoris Komagataella Lamiales Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaffomycetaceae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Rhabdoviridae Riboviria Ricinus communis Ricinus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Schizotrypanum Simiiformes Sophophora Spermatophyta Spinacia oleracea Spinacia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified Vesiculovirus unidentified influenza virus
G86377KB	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex Sda/CT antigen	Linkage defined saccharide	4290.53	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86375CM	Fuc(a1-2)Gal(b1-?)[Fuc(a1-?)]GlcNAc(b1-3)Gal(b1-3)GalNAc(a1-	Blood group H (type 1) Blood group H (type 2) Blood group H (type 2) Leb Leb Lewis b Lewis b Ley O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	1040.39	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86372GF	Gal(b1-3)[Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)GlcNAc(b1-6)]GalNAc(?1-	O-Glycan core 2 fuzzy Sda/CT antigen	Linkage defined saccharide	1242.45	Amniota Bilateria Boreoeutheria Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Macaca radiata Macaca Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86357DX	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2151.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86355AM			Linkage defined saccharide	400.16	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86343GW	Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	I' antigen N-Glycan complex Type 2 LN2	Linkage defined saccharide	4153.48	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86321EF			Linkage defined saccharide	1052.03	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86298IX	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	Blood group A (type 2) N-Glycan complex Polysialic acid	Linkage defined saccharide	2426.88	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86297CD	Fuc(a1-4)[Fuc(a1-5)]Kdn(a2-3)Gal(b1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)]GalNAc-ol	Blood group H (type 2) Leb Lewis b Ley	Linkage defined saccharide	1584.59	Ambystoma tigrinum Ambystoma Ambystomatidae Amphibia Batrachia Bilateria Caudata Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Opisthokonta Salamandroidea Sarcopterygii Teleostomi Tetrapoda Vertebrata cellular organisms