GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1876 - 1900** of **5866** in total

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GlyTouCan ID	IUPAC Condensed	Motifs ▲	Subsumption Level	Monoisotopic Mass	Organisms
G23901NY	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Fuc(a1-2)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24202BK		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2280.82	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24585JV	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1769.63	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Perissodactyla Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G24604HV	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2005.72	Alphavirus Alsuviricetes Amniota Archelosauria Archosauria Artiodactyla Aves Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Laurasiatheria Mammalia Martellivirales Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Orthornavirae Pecora Phasianidae Phasianinae Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Semliki Forest virus Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Togaviridae Torpedinidae Torpediniformes Vertebrata Viruses artificial sequences cellular organisms other entries other sequences synthetic construct unclassified entries unclassified sequences unidentified
G24501HF		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2661.95	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G25140TA	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2134.77	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G25837HW	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[Gal(b1-4)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2077.75	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G25520XG	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Archelosauria Archosauria Articulavirales Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Perdicinae Phasianidae Polyploviricotina Primates Rattus norvegicus Rattus Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G25481DT		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1681.62	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G26145IL		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G25713RA		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2733.97	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G26730QM	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-4)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2735.99	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G26877ST	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2222.78	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G27919IH	GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1624.6	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G28106CM	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	3390.2	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G28156XV	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(?1-?)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G28459KM	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2879.01	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G28659UB	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2208.8	Amniota Archelosauria Archosauria Aves Batoidea Bilateria Boreoeutheria Catarrhini Chondrichthyes Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Meleagridinae Meleagris gallopavo Meleagris Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Vertebrata cellular organisms
G29024OJ	GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1624.6	Amniota Archelosauria Archosauria Articulavirales Artiodactyla Aves Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Capra hircus Capra Caprinae Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Chondrichthyes Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insthoviricetes Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Perissodactyla Phasianidae Phasianinae Polyploviricotina Primates Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Vertebrata Viruses artificial sequences cellular organisms other entries other sequences synthetic construct unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G28362DW		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2077.75	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G29972ZX	GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1681.62	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G29857RC	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2005.72	Alphavirus Alsuviricetes Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Laurasiatheria Mammalia Martellivirales Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Orthornavirae Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Semliki Forest virus Simiiformes Teleostomi Tetrapoda Theria Theropoda Togaviridae Vertebrata Viruses cellular organisms
G29228MX	Gal(b1-4)GlcNAc(b1-?)[GlcNAc(b1-?)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1989.73	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G29884LW	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2280.82	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G30529FG	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2005.72	Acomys russatus Acomys Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deomyinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms