GlyCosmos Portal

Table Filtering

GlyTouCan ID

IUPAC Condensed

Motifs

Subsumption Level

Monoisotopic Mass

Organisms

Taxonomy IDs

Tissues

Display
results per page.

Clear

Submissions

GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **2051 - 2075** of **9190** in total

« First

‹ Prev

…

79

80

81

82

83

84

85

86

87

…

Next ›

Last »
GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G99182WQ	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	I' antigen N-Glycan complex Type 2 LN2	Linkage defined saccharide	2370.86	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G43669FQ			Base composition with linkage	2370.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G50535IT	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3/6)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64841EF	Gal(b1-?)GlcNAc(b1-2)[Gal(b1-?)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-?)GlcNAc(b1-2)[Gal(b1-?)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 1 LN Type 2 LN	Linkage defined saccharide	2370.86	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88800UV	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G11561RV	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	Blood group H (type 2) Leb Lewis b Ley N-Glycan complex Polysialic acid	Linkage defined saccharide	2369.86	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G51413EV			Glycosidic topology	2369.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G65635AB	Neu5Ac(a2-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Sialyl Lewis x Slex	Linkage defined saccharide	2369.86	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G07799LX			Monosaccharide composition with linkage	2369.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G00318AV	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69725KG	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Perissodactyla Primates Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G91060IP	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Neu5Ac(?2-?)Gal(?1-?)[Fuc(a1-3)]GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G72954XC	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2368.84	50 kb inversion clade Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Butea monosperma Butea Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neognathae Opisthokonta Papilionoideae Pecora Pentapetalae Perissodactyla Phaseoleae Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids unclassified entries unclassified sequences unidentified
G90889KG	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Actinopteri Actinopterygii Alphavirus Alsuviricetes Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops aethiops Chlorocebus aethiops Chlorocebus Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Laurasiatheria Mammalia Martellivirales Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Orthornavirae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Sindbis virus Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms unclassified entries unclassified sequences unidentified
G04286YE	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2/4)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2/4)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G22836DF	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G24005JZ	Neu5Ac(a2-3)Gal(?1-?)GlcNAc(?1-?)Man(a1-3/6)[Neu5Ac(a2-3)Gal(?1-?)GlcNAc(?1-?)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G34946TH	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G35242IF	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04437UY	Neu5Ac(a2-?)Hex(?1-?)HexNAc(?1-?)Hex(?1-?)[Neu5Ac(a2-?)Hex(?1-?)HexNAc(?1-?)Hex(?1-?)]Hex(?1-?)HexNAc(?1-?)[Fuc(?1-?)]HexNAc(b1-		Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05610JO		N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G07097VH		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G08146BT	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2368.84	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G08606CV	Neu5Ac(a2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Neu5Ac(a2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2368.84	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G14899VS	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	2368.84	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms