GlyCosmos Portal

Source	Last Updated
GlyTouCan	January 5, 2026

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GlyTouCan ID	IUPAC Condensed	Names ▲	Motifs	Archetype	Subsumption Level	Monoisotopic Mass	Organisms
G19059PI		Chondroitin-4-Sulfate		G52135DT	Fully-defined saccharide		Acalyptratae Actinopteri Actinopterygii Amniota Ampullariidae Ampullarioidea Archelosauria Architaenioglossa Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenogastropoda Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Gerbillinae Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Lagomorpha Laurasiatheria Leporidae Lophotrochozoa Macaca mulatta Macaca Mammalia Mandibulata Meriones unguiculatus Meriones Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Pomacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spiralia Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Whippomorpha cellular organisms melanogaster group melanogaster subgroup
G01614ZM	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	Disialyl T Antigen	Di-sialyl T antigen	G30110UW	Fully-defined saccharide	965.33	Actinopteri Actinopterygii Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend murine leukemia virus Gammaretrovirus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ortervirales Orthoretrovirinae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pararnavirae Pecora Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G46677TE	Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1a	GD1a	G21518JT	Fully-defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carangaria Carassius auratus Carassius Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chinchillidae Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Delphinidae Deuterostomia Dinosauria Dipnotetrapodomorpha Discoba Euacanthomorphacea Euarchontoglires Euglenozoa Eukaryota Eulipotyphla Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Kinetoplastea Lagomorpha Laurasiatheria Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Leporidae Mammalia Meleagridinae Meleagris gallopavo Meleagris Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Odontoceti Oncorhynchus mykiss Oncorhynchus Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Papio hamadryas Papio Parophrys vetulus Parophrys Pecora Percomorphaceae Phasianidae Phasianinae Pleuronectidae Pleuronectiformes Pleuronectoidei Primates Protacanthopterygii Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Soricidae Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trypanosomatida Trypanosomatidae Tursiops truncatus Tursiops Vertebrata Whippomorpha cellular organisms
G37184KW	Neu5Ac(a2-8)Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD1b	GD1b	G84052PW	Fully-defined saccharide	1289.44	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Channichthyidae Chionodraco myersi Chionodraco Chondrichthyes Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Crocidurinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Euacanthomorphacea Euarchontoglires Eukaryota Eulipotyphla Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Leucoraja erinacea Leucoraja Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Narcinidae Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Papio hamadryas Papio Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Pipidae Pipoidea Primates Pseudocrenilabrinae Rajidae Rajiformes Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Soricidae Stenella coeruleoalba Stenella Suidae Suina Suncus murinus Suncus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Xenopodinae Xenopus laevis Xenopus Xenopus Zeiogadaria cellular organisms
G02657AK	Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GD2	GD2	G98397EB	Fully-defined saccharide	1127.39	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Pipidae Pipoidea Primates Pseudocrenilabrinae Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Vertebrata Xenopodinae Xenopus cellular organisms
G98544DH	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	GD3 N-Acetyl GD3	9-O-Acetyl GD3 GD3	G10158SF	Fully-defined saccharide	924.31	Actinopteri Actinopterygii Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Balaenoptera edeni Balaenoptera Balaenopteridae Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chinchilla Chinchillidae Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leucoraja erinacea Leucoraja Macaca fuscata Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Narcinidae Neognathae Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Papio hamadryas Papio Pecora Phasianidae Phasianinae Pipidae Pipoidea Primates Rajidae Rajiformes Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Vertebrata Whippomorpha Xenopodinae Xenopus cellular organisms
G48558GR	Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM1 GM1a	GM1	G41876BL	Fully-defined saccharide	998.34	Actinopteri Actinopterygii Bacteria Bilateria Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Carassius auratus Carassius Chordata Clupeocephala Craniata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Discoba Epsilonproteobacteria Euglenozoa Eukaryota Eumetazoa Euteleostomi Gnathostomata Kinetoplastea Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Metakinetoplastina Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Trypanosomatida Trypanosomatidae Vertebrata cellular organisms
G79389NT	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM2 N-Acetyl GM2	GM2	G65156XZ	Fully-defined saccharide	836.29	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Carassius auratus Carassius Catarrhini Chordata Clupeocephala Craniata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Protacanthopterygii Salmonidae Salmoniformes Salmoninae Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G91237TK	Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	GM3 N-Acetyl GM3	GM3	G40764VP	Fully-defined saccharide	633.21	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Channichthyidae Chinchilla Chinchillidae Chionodraco myersi Chionodraco Chondrichthyes Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Enterobacterales Enterobacteriaceae Epsilonproteobacteria Escherichia coli Escherichia Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Leucoraja erinacea Leucoraja Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Narcinidae Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Oncorhynchus mykiss Oncorhynchus Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Ovis aries Ovis Papio hamadryas Papio Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Pipidae Pipoidea Platyrrhini Primates Protacanthopterygii Pseudocrenilabrinae Pseudomonadota Rajidae Rajiformes Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Trematomus lepidorhinus Trematomus Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus Zeiogadaria cellular organisms
G30207PZ	Neu5Ac(a2-3)Gal(b1-	GM4 Neu5Ac(a2-3)Gal(b1)	GM4	G58436AI	Fully-defined saccharide	471.16	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cetacea Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Zeiogadaria cellular organisms
G26203DP	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GP1c	GQ1b GQ1c	G44362TS	Fully-defined saccharide	2162.73	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Carassius auratus Carassius Catarrhini Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neopterygii Neoteleostei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Percomorphaceae Phasianidae Phasianinae Primates Pseudocrenilabrinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G18625KA	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GQ1b	GQ1b	G71158TZ	Fully-defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Carassius auratus Carassius Carnivora Catarrhini Cetacea Channichthyidae Chinchilla Chinchillidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Epsilonproteobacteria Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha cellular organisms
G68110IF	Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1a	GT1a	G97898ZO	Fully-defined saccharide	1580.53	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Bilateria Boreoeutheria Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Catarrhini Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Epsilonproteobacteria Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Primates Pseudocrenilabrinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms
G40183QN	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT1b	GT1b	G08648UJ	Fully-defined saccharide	1580.53	Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Carassius auratus Carassius Chordata Clupeocephala Coelurosauria Craniata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Meleagridinae Meleagris gallopavo Meleagris Metazoa Neognathae Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G36476BA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT2	GT2	G66281XG	Fully-defined saccharide	1418.48	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Artiodactyla Bilateria Boreoeutheria Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus morhua Gadus Glires Gnathostomata Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muroidea Myomorpha Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phodopus sungorus Phodopus Pollachius pollachius Pollachius Pseudocrenilabrinae Rodentia Sarcopterygii Scardinius erythrophthalmus Scardinius Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms
G56516VH	Neu5Ac(a2-3)Gal(b1-4)Glc(?1-	Neu5Ac(a2-3)Gal(b1-4)Glc	GM4	G40764VP	Linkage-defined saccharide	633.21	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G81928IQ	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(?1-	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc	GM4	G10158SF	Linkage-defined saccharide	924.31	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G53517XC	Neu5Gc(a2-3)Gal(?1-	Neu5Gc(a2-3)Gal		G18324XR	Linkage-defined saccharide	487.15	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Perissodactyla Sarcopterygii Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10109RQ	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[Gal(b1-4)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-		N-Glycan complex Polysialic acid	G45411YF	Linkage-defined saccharide	2077.75	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G24439WH	Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		N-Glycan complex N-acetyllactosamine Type 2 LN	G73044VU	Linkage-defined saccharide	1478.54	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G48209XU	Kdn(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[GlcNAc(b1-2)Man(a1-?)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-		N-Glycan complex N-acetyllactosamine Type 2 LN	G41276XE	Linkage-defined saccharide	2077.75	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G88384ML	Neu5Ac(a2-3)[Gal(b1-4)]Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)[Gal(b1-4)]Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc		Polysialic acid		Linkage-defined saccharide	2343.81	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Leuciscidae Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pseudaspininae Pseudaspius hakonensis Pseudaspius Teleostei Teleostomi Vertebrata cellular organisms
G06123VV	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		N-Glycan complex Polysialic acid	G78291UW	Linkage-defined saccharide	2093.74	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G14633CL	Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc(?1-		GM4	G01269WV	Linkage-defined saccharide	998.34	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G77103HZ	Fuc(a1-?)Fuc(a1-3)HexNAc(?1-?)HexNAc(?1-?)Hex(?1-?)Hex(?1-?)Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(?1-		N-acetyllactosamine Type 2 LN a-Gal antigen	G84961RY	Linkage-defined saccharide	2094.76	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms

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