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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 4, 2025

Displaying entries **1 - 25** of 41 in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Archetype	Subsumption Level	Monoisotopic Mass	Organisms
G02524YV	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc			Linkage-defined saccharide	1154.43	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G05854FB	Gal(a1-3)[GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-acetyllactosamine Type 2 LN a-Gal antigen	G87111OL	Linkage-defined saccharide	1640.59	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G12447LD	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-3)Gal(b1-4)Glc(b1-	GM4 Lactosylceramide		Fully-defined saccharide	1248.41	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Poeciliidae Poeciliinae Teleostei Teleostomi Vertebrata Xiphophorus cellular organisms
G14745ZW	Gal(a1-3)[Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-acetyllactosamine Type 2 LN a-Gal antigen		Fully-defined saccharide	1437.51	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G27415TQ	Gal(b1-4)Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-acetyllactosamine Type 2 LN	G17494WZ	Linkage-defined saccharide	1275.46	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G28952ZD	Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)][Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	G40171MO	Linkage-defined saccharide	5291.94	Acanthomorphata Actinopteri Actinopterygii Bilateria Carangaria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paralichthyidae Paralichthys olivaceus Paralichthys Percomorphaceae Pleuronectiformes Pleuronectoidei Teleostei Teleostomi Vertebrata cellular organisms
G30269YQ	Neu5Ac(a2-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	GM4 N-Glycan complex N-acetyllactosamine Type 2 LN		Linkage-defined saccharide	2546.89	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias latipes Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G36476BA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GT2	G66281XG	Fully-defined saccharide	1418.48	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Artiodactyla Bilateria Boreoeutheria Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus morhua Gadus Glires Gnathostomata Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muroidea Myomorpha Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Perciformes Percomorphaceae Phodopus sungorus Phodopus Pollachius pollachius Pollachius Pseudocrenilabrinae Rodentia Sarcopterygii Scardinius erythrophthalmus Scardinius Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Trematomus lepidorhinus Trematomus Vertebrata Zeiogadaria cellular organisms
G38424XU	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		G02524YV	Linkage-defined saccharide	1154.43	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G95493MF	Neu5Ac(a2-3)[GalNAc(a1-3)GalNAc(b1-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM1	G05049TB	Fully-defined saccharide	1404.5	Acanthomorphata Actinopteri Actinopterygii Bilateria Carangaria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Parophrys vetulus Parophrys Percomorphaceae Pleuronectidae Pleuronectiformes Pleuronectoidei Teleostei Teleostomi Vertebrata cellular organisms
G99524KA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GQ1c	G54434YJ	Fully-defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Phasianidae Phasianinae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G13422DI	Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)][Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	G40171MO	Fully-defined saccharide	5291.94	Acanthomorphata Actinopteri Actinopterygii Bilateria Carangaria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Percomorphaceae Pleuronectiformes Pleuronectoidei Teleostei Teleostomi Vertebrata cellular organisms
G17907WL	Gal(a1-3)Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-acetyllactosamine Type 2 LN a-Gal antigen	G74946ZV	Linkage-defined saccharide	1275.46	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G28431AN		I' antigen Type 2 LN2		Linkage-defined saccharide		Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G32534WD		N-Glycan complex SDLex		Fully-defined saccharide		Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G38697HB	GalNAc(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	LDN LacdiNAc		Fully-defined saccharide	1154.43	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G38941QE	Gal(a1-3)Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-acetyllactosamine Type 2 LN a-Gal antigen		Fully-defined saccharide	1275.46	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G59965VP	Gal(a1-3)[GalNAc(b1-4)GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-acetyllactosamine Type 2 LN a-Gal antigen	G23489QR	Linkage-defined saccharide	1843.67	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G60384JB	Gal(a1-3)[Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-acetyllactosamine Type 2 LN a-Gal antigen	G48285QA	Linkage-defined saccharide	1437.51	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G63899VB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex		Linkage-defined saccharide	1722.65	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G77337WY	Gal(a1-3)[GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-acetyllactosamine Type 2 LN a-Gal antigen		Fully-defined saccharide	1640.59	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G78553RL	Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)][Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-3)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Fuc(a1-3)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	G48632WP	Linkage-defined saccharide	5291.94	Acanthomorphata Actinopteri Actinopterygii Bilateria Carangaria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paralichthyidae Paralichthys olivaceus Paralichthys Percomorphaceae Pleuronectiformes Pleuronectoidei Teleostei Teleostomi Vertebrata cellular organisms
G84319PN	Neu5Ac(a2-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	GM4 N-Glycan complex N-acetyllactosamine Type 2 LN	G30269YQ	Linkage-defined saccharide	2546.89	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias latipes Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G95978MK	Gal(b1-4)Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-acetyllactosamine Type 2 LN		Fully-defined saccharide	1275.46	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G95920TH	Gal(a1-3)[GalNAc(b1-4)GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	LDN LacdiNAc N-acetyllactosamine Type 2 LN a-Gal antigen		Fully-defined saccharide	1843.67	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms